A revised classification of the family Dasyatidae (Chondrichthyes: Myliobatiformes) based on new morphological and molecular insights Author Last, Peter R. Author Naylor, Gavin J. P. Author Manjaji-Matsumoto, B. Mabel text Zootaxa 2016 4139 3 345 368 journal article 10.11646/zootaxa.4139.3.2 51358935-98b0-4d89-a47f-e4341171e5b9 1175-5326 262765 DCF4220B-4A73-407C-837C-54DEAE29F435 Genus Makararaja Roberts, 2007 Makararaja Roberts, 2007 :286 . Type species Makararaja chindwinensis Roberts 2007 ; by original designation and monotypic. Definition . Small dasyatids (to at least 39 cm DW) characterised by the following: relatively well-depressed, oval to almost circular disc with pectoral-fin apex broadly rounded; snout obtuse and medium length (~1.7 times combined orbit and spiracle length); eye small and protruding slightly; nasal curtain distinctly bilobed; mouth narrow with 4 oral papillae; tail flexible and short (length ~1.5 times DW), its base moderately broad and depressed; pelvic fins large, protruding slightly beyond disc; dorsal fold absent; ventral fold moderately deep with a short base; caudal sting positioned very posteriorly on tail (distance from pectoral-fin insertion to caudal-sting base ~5.3 times interspiracular width); skin on dorsal surface uniformly velvety, densely covered with minute denticles and lacking a denticle band; no enlarged thorns on disc or tail (but with a short nuchal row of rudimentary thorns); plain coloured dorsally; freshwater, South-East Asia. Species . M. chindwinensis Roberts, 2007 Remarks . Poorly known (based on the holotype ); more specimens and tissues needed.