Molecular and morphological characterisation of new species in the trapdoor spider genus Aname (Araneae: Mygalomorphae: Nemesiidae) from the Pilbara bioregion of Western Australia
Author
Harvey, Frances S. B.
Author
Framenau, Volker W.
Author
Wojcieszek, Janine M.
Author
Rix, Michael G.
Author
Harvey, Mark S.
text
Zootaxa
2012
3383
15
38
journal article
10.5281/zenodo.215484
1b3a0373-3529-4147-b69b-4f4d9d9e5804
1175-5326
215484
Genus
Aname
L. Koch, 1873
Aname
L. Koch 1873
: 465
.
Type
species
Aname pallida
L. Koch, 1873
, by monotypy.
Dekana
Hogg 1902
: 138
(synonymised by
Raven 1981
: 328
and
Main 1982b
: 27
).
Type
species
Dekana diversicolor
Hogg, 1902
, by original designation.
Proshermacha
Simon 1908
: 113
(synonymised with
Chenistonia
by
Main 1982a
: 113
).
Type
species
Proshermacha subarmata
Simon, 1908
(junior synonym of
Chenistonia teppperi
Hogg, 1902
), by subsequent designation of
Rainbow (1911)
.
Sungenia
Rainbow and Pulleine 1918
: 162
(synonymised by
Raven 1981
: 328
and
Main 1982b
: 27
).
Type
species
Chenistonia
(
Dekana
)
atra
Strand, 1913
, by monotypy.
Dolichosternum
Rainbow and Pulleine 1918
: 168
(synonymised by
Raven 1981
: 328
).
Type
species
Dolichosternum attenuatum
Rainbow and Pulleine, 1918
(junior synonym of
Ixamatus distinctus
Rainbow, 1914
), by monotypy.
Diagnosis.
Species of
Aname
differ from all other nemesiids by the presence of a large mega-spur on the tibia of leg I generally in a mid-distal position (e.g.
Fig. 15
), the absence of spines on the pedal tarsi (e.g.
Fig. 19
), and the absence of cuspules on the pedal coxae (e.g.
Fig. 12
) (following
Main 1982b
,
1983
,
1986
;
Raven 1981
). The male pedipalpal bulb is spherical to pear-shaped, with the embolus originating from the bulb in a distal position (e.g.
Figs 17–20
); the shape of the embolus is usually long and slender, and slightly curved in profile (e.g.
Figs 17–20
).
Remarks
. While we are confident that there are four distinct species represented in our samples, we found considerable difficulty in distinguishing these species from some previously named species. Differentiating our species from those for which males have been described was not difficult. Several species of
Aname
are currently known only from adult females and for which spermathecae have not been described or illustrated. These are:
A. armigera
Rainbow & Pulleine, 1918
(from south-western
Australia
),
A. aurea
Rainbow & Pulleine, 1918
(from New South
Wales
),
A. coenosa
Rainbow & Pulleine, 1918
(from South
Australia
),
A. comosa
Rainbow & Pulleine, 1918
(from South
Australia
),
A. cuspidata
(
Main, 1954
)
(from south-western
Australia
),
A. fuscocincta
Rainbow & Pulleine, 1918
(from south-western
Australia
),
A. grandis
Rainbow & Pulleine, 1918
(from South
Australia
),
A. hirsuta
Rainbow & Pulleine, 1918
(from South
Australia
),
A. maculata
(
Rainbow & Pulleine, 1918
)
(from south-western
Australia
),
A. platypus
(
L. Koch, 1875
)
(possibly from
Australia
),
A. tasmanica
Hogg, 1902
(from Tasmania) and
A. villosa
(
Rainbow & Pulleine, 1918
)
(from south-western
Australia
) (
Hogg 1902
;
Koch 1875
;
Main 1954
;
Rainbow & Pulleine 1918
).
As
the
type
localities of all of these species are located at least
1,000 km
away from the Pilbara region, we are confident that we have not inadvertently described a species that will eventually become a junior synonym.