<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.126.34096" ID-PMC="PMC6650443" ID-Pensoft-Pub="1314-2003-126-89" ID-Pensoft-UUID="FFC2D06D486FF317CE32972BDE2BFF93" ID-PubMed="31360096" ID-Zenodo-Dep="3348547" ModsDocID="1314-2003-126-89" checkinTime="1563287727035" checkinUser="pensoft" docAuthor="Peterson, Paul M., Romaschenko, Konstantin, Soreng, Robert J. &amp; Reyna, Jesus Valdes" docDate="2019" docId="184BEF72E4915F5E8442F5ED837EAD73" docLanguage="en" docName="PhytoKeys 126: 89-125" docOrigin="PhytoKeys 126" docPubDate="2019-07-16" docSource="http://dx.doi.org/10.3897/phytokeys.126.34096" docTitle="Barkworthia stillmanii (Bol.) Romasch., P. M. Peterson &amp; Soreng, comb. nov." docType="treatment" docVersion="3" id="FFC2D06D486FF317CE32972BDE2BFF93" lastPageNumber="93" masterDocId="FFC2D06D486FF317CE32972BDE2BFF93" masterDocTitle="A key to the North American genera of Stipeae (Poaceae, Pooideae) with descriptions and taxonomic names for species of Eriocoma, Neotrinia, Oloptum, and five new genera: Barkworthia, x Eriosella, Pseudoeriocoma, Ptilagrostiella, and Thorneochloa" masterLastPageNumber="125" masterPageNumber="89" pageNumber="92" updateTime="1668139369136" updateUser="ExternalLinkService">
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<mods:title>A key to the North American genera of Stipeae (Poaceae, Pooideae) with descriptions and taxonomic names for species of Eriocoma, Neotrinia, Oloptum, and five new genera: Barkworthia, x Eriosella, Pseudoeriocoma, Ptilagrostiella, and Thorneochloa</mods:title>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Peterson, Paul M.</mods:namePart>
<mods:affiliation>Department of Botany MRC- 166, National Museum of Natural History, Smithsonian Institution, Washington, DC 20013 - 7012, USA</mods:affiliation>
<mods:nameIdentifier type="email">peterson@si.edu</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Romaschenko, Konstantin</mods:namePart>
<mods:affiliation>Department of Botany MRC- 166, National Museum of Natural History, Smithsonian Institution, Washington, DC 20013 - 7012, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Soreng, Robert J.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-8358-4915</mods:nameIdentifier>
<mods:affiliation>Department of Botany MRC- 166, National Museum of Natural History, Smithsonian Institution, Washington, DC 20013 - 7012, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Reyna, Jesus Valdes</mods:namePart>
<mods:affiliation>Departamento de Botanica, Universidad Autonoma Agraria Antonio Narro, Saltillo, C. P. 25315, Mexico</mods:affiliation>
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<mods:title>PhytoKeys</mods:title>
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<mods:date>2019</mods:date>
<mods:detail type="pubDate">
<mods:number>2019-07-16</mods:number>
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<mods:detail type="volume">
<mods:number>126</mods:number>
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<mods:start>89</mods:start>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.126.34096</mods:url>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.126.34096</mods:identifier>
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<paragraph pageId="3" pageNumber="92">
<taxonomicName LSID="184BEF72-E491-5F5E-8442-F5ED837EAD73" authority="(Bol.) Romasch., P. M. Peterson &amp; Soreng, comb. nov." authorityName="(Bol.) Romasch., P. M. Peterson &amp; Soreng, comb. nov." class="Liliopsida" family="Poaceae" genus="Barkworthia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Barkworthia stillmanii" order="Poales" pageId="3" pageNumber="92" phylum="Tracheophyta" rank="species" species="stillmanii">Barkworthia stillmanii (Bol.) Romasch., P.M.Peterson &amp; Soreng, comb. nov.</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="92" type="reference_group">
<paragraph pageId="3" pageNumber="92">
<taxonomicName authorityName="Bol" authorityYear="1872" class="Liliopsida" family="Poaceae" genus="Stipa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Stipa stillmanii" order="Poales" pageId="3" pageNumber="92" phylum="Tracheophyta" rank="species" species="stillmanii">Stipa stillmanii</taxonomicName>
Bol., Proc. Calif. Acad. Sci. 4: 169. 1872 [Basionym] ≡ 
<taxonomicName class="Liliopsida" family="Poaceae" genus="Achnatherum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Achnatherum stillmanii" order="Poales" pageId="3" pageNumber="92" phylum="Tracheophyta" rank="species" species="stillmanii">Achnatherum stillmanii</taxonomicName>
(Bol.) Barkworth, Phytologia 74(1): 14. 1993 - Type: USA, California, Sierra Nevada, Blue 
<normalizedToken originalValue="Cañon">Canon</normalizedToken>
, Jul 1870, 
<emphasis italics="true" pageId="3" pageNumber="92">H.N. Bolander</emphasis>
, 
<emphasis italics="true" pageId="3" pageNumber="92">M.D. Kellogg &amp; co. s.n.</emphasis>
(holotype: NY-00431576 [image!]; isotypes: GH-00017890 [image!], K-000873398 [image!], MO-3055652!, MO-3055653!, MO-3055654!, NDG-07159 [image!], US-556922!). Fig. 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Barkworthia stillmanii: A habit B ligule C panicle D glumes E floret F floret (close up). Eriocoma hymenoides: G culm and panicle H panicle branch I glumes J floret K lemma awn L floret base (callus)." figureDoi="10.3897/phytokeys.126.34096.figure1" httpUri="https://binary.pensoft.net/fig/317891" pageId="3" pageNumber="92">1A-F</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="92" type="distribution">
<paragraph pageId="3" pageNumber="92">Distribution and habitat.</paragraph>
<paragraph pageId="3" pageNumber="92">
<taxonomicName class="Liliopsida" family="Poaceae" genus="Barkworthia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Barkworthia stillmanii" order="Poales" pageId="3" pageNumber="92" phylum="Tracheophyta" rank="species" species="stillmanii">
<emphasis italics="true" pageId="3" pageNumber="92">Barkworthia stillmanii</emphasis>
</taxonomicName>
is distributed in scattered locations in northern California (Butte, El Dorado, Nevada, Placer, Plumas, Sacramento, Shasta, Sierra, Tehama, Trinity, Tulare, and Yuba Counties) associated with yellow pine and red fir forests; 10-1500 m (
<bibRefCitation author="Barkworth, ME" editor="Barkworth, ME" journalOrPublisher="Oxford Universtiy Press, New York" pageId="20" pageNumber="109" pagination="109 - 186" refId="B3" refString="Barkworth, ME, 2007. 10. Stipeae Dumort. In: Barkworth, ME, Capels, KM, Long, S, Anderton, LK, Piep, MB, Eds., Magnoliophyta: Commelinidae (in part): Poaceae, part 1. Oxford Universtiy Press, New York: 109 - 186" title="10. Stipeae Dumort." volumeTitle="Magnoliophyta: Commelinidae (in part): Poaceae, part 1." year="2007">Barkworth 2007</bibRefCitation>
; 
<bibRefCitation author="Calflora" journalOrPublisher="Smithsonian Institution Press, Washington, D. C." pageId="20" pageNumber="109" publicationUrl="https://www.calflora.org/" refId="B5" refString="Calflora, 2018. Information on California plants for education, research and conservation. Berkeley, California: The Calflora Database (a non-profit organization). https://www.calflora.org/" title="Information on California plants for education, research and conservation. Berkeley, California: The Calflora Database (a non-profit organization)." url="https://www.calflora.org/" year="2018">Calflora 2018</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="93" type="comments">
<paragraph pageId="4" pageNumber="93">
<pageBreakToken pageId="4" pageNumber="93" start="start">Comments</pageBreakToken>
.
</paragraph>
<paragraph pageId="4" pageNumber="93">
In a molecular-derived phylogeny of the 
<taxonomicName family="Poaceae" kingdom="Plantae" lsidName="" order="Poales" pageId="4" pageNumber="93" rank="tribe" tribe="Stipeae">Stipeae</taxonomicName>
using 10 DNA markers 
<taxonomicName class="Liliopsida" family="Poaceae" genus="Barkworthia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Barkworthia stillmanii" order="Poales" pageId="4" pageNumber="93" phylum="Tracheophyta" rank="species" species="stillmanii">
<emphasis italics="true" pageId="4" pageNumber="93">Barkworthia stillmanii</emphasis>
</taxonomicName>
is sister to a well-supported 
<taxonomicName class="Liliopsida" family="Gramineae" genus="Piptatherum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Piptatherum" order="Poales" pageId="4" pageNumber="93" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="4" pageNumber="93">Piptatherum</emphasis>
</taxonomicName>
clade, which is strictly Old World in distribution, and has cauducous awns, and dark glossy lemmas in fruit (
<bibRefCitation DOI="https://doi.org/10.1002/tax.611002" author="Romaschenko, K" journalOrPublisher="Taxon" pageId="22" pageNumber="111" pagination="18 - 44" refId="B26" refString="Romaschenko, K, Peterson, PM, Soreng, RJ, Garcia-Jacas, N, Futorna, O, Susanna, A, 2012. Systematics and evolution of the needle grasses (Poaceae: Pooideae: Stipeae) based on analysis of multiple chloroplast loci, ITS, and lemma micromorphology. Taxon 61 (1): 18 - 44, DOI: https://doi.org/10.1002/tax.611002" title="Systematics and evolution of the needle grasses (Poaceae: Pooideae: Stipeae) based on analysis of multiple chloroplast loci, ITS, and lemma micromorphology." url="https://doi.org/10.1002/tax.611002" volume="61" year="2012">Romaschenko et al. 2012</bibRefCitation>
).
</paragraph>
<caption ID-Zenodo-Dep="3348567" doi="10.3897/phytokeys.126.34096.figure1" httpUri="https://binary.pensoft.net/fig/317891" pageId="4" pageNumber="93" start="Figure 1" startId="F1">
<paragraph pageId="4" pageNumber="93">
<emphasis bold="true" pageId="4" pageNumber="93">Figure 1.</emphasis>
<taxonomicName class="Liliopsida" family="Poaceae" genus="Barkworthia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Barkworthia stillmanii" order="Poales" pageId="4" pageNumber="93" phylum="Tracheophyta" rank="species" species="stillmanii">
<emphasis italics="true" pageId="4" pageNumber="93">Barkworthia stillmanii</emphasis>
</taxonomicName>
: 
<emphasis bold="true" pageId="4" pageNumber="93">A</emphasis>
habit 
<emphasis bold="true" pageId="4" pageNumber="93">B</emphasis>
ligule 
<emphasis bold="true" pageId="4" pageNumber="93">C</emphasis>
panicle 
<emphasis bold="true" pageId="4" pageNumber="93">D</emphasis>
glumes 
<emphasis bold="true" pageId="4" pageNumber="93">E</emphasis>
floret 
<emphasis bold="true" pageId="4" pageNumber="93">F</emphasis>
floret (close up). 
<taxonomicName class="Liliopsida" family="Poaceae" genus="Eriocoma" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Eriocoma hymenoides" order="Poales" pageId="4" pageNumber="93" phylum="Tracheophyta" rank="species" species="hymenoides">
<emphasis italics="true" pageId="4" pageNumber="93">Eriocoma hymenoides</emphasis>
</taxonomicName>
: 
<emphasis bold="true" pageId="4" pageNumber="93">G</emphasis>
culm and panicle 
<emphasis bold="true" pageId="4" pageNumber="93">H</emphasis>
panicle branch 
<emphasis bold="true" pageId="4" pageNumber="93">I</emphasis>
glumes 
<emphasis bold="true" pageId="4" pageNumber="93">J</emphasis>
floret 
<emphasis bold="true" pageId="4" pageNumber="93">K</emphasis>
lemma awn 
<emphasis bold="true" pageId="4" pageNumber="93">L</emphasis>
floret base (callus).
</paragraph>
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