<document id="A46E1FCBB6B5327AD382AF22F9431BD7" ID-DOI="10.11646/zootaxa.5481.3.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="12758944" ID-ZooBank="F86F4482-5F80-444F-A5DE-F1F1BAC04E41" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721216078619" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2024" docId="A64A2B29256A163142FEFE5EFD669E2F" docLanguage="en" docName="zootaxa.5481.3.7.pdf" docOrigin="Zootaxa 5481 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5481.3.7" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hygrodromicus coriaceus" docType="treatment" docVersion="3" lastPageNumber="385" masterDocId="5A735351256B16304269FFD9FFC49D3C" masterDocTitle="On some species of the genus Hygrodromicus Tronquet with description of a new species from Pakistan (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)" masterLastPageNumber="390" masterPageNumber="384" pageNumber="385" updateTime="1721216879074" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="D8765351A42537AF681AE76AE040EDBF">On some species of the genus Hygrodromicus Tronquet with description of a new species from Pakistan (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)</mods:title>
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<taxonomicName id="E9E3E1BC256A163142FEFE5EFD6C9C9E" ID-CoL="3NBHT" authority="(Cameron, 1924)" baseAuthorityName="Cameron" baseAuthorityYear="1924" box="[151,680,391,418]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="385" phylum="Arthropoda" rank="species" species="coriaceus">
<emphasis id="1C97462D256A163142FEFE5EFD6C9C9E" bold="true" box="[151,680,391,418]" pageId="1" pageNumber="385">
<emphasis id="1C97462D256A163142FEFE5EFE149C9E" bold="true" box="[151,464,391,418]" italics="true" pageId="1" pageNumber="385">Hygrodromicus coriaceus</emphasis>
(
<bibRefCitation id="4A72E7CE256A163143B6FE51FD649C9E" author="Cameron, M." box="[479,672,392,418]" pageId="1" pageNumber="385" pagination="160 - 198" refId="ref3298" refString="Cameron, M. (1924) New species of Staphylinidae from India. Transactions of the Entomological Society of London, 1924, 160 - 198. https: // doi. org / 10.1111 / j. 1365 - 2311.1924. tb 03357. x" type="journal article" year="1924">Cameron, 1924</bibRefCitation>
)
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</heading>
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<subSubSection id="66F9C9B4256A163142FEFE15FC5B9F63" pageId="1" pageNumber="385" type="reference_group">
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<treatmentCitation id="AF42BC2E256A163142FEFE15FDA49CD9" author="Cameron, M." box="[151,608,460,485]" page="172" pageId="1" pageNumber="385" year="1924">
<taxonomicName id="E9E3E1BC256A163142FEFE15FDA49CD9" ID-CoL="3FPLC" authority="Cameron, 1924: 172" authorityName="Cameron" authorityPageNumber="172" authorityYear="1924" box="[151,608,460,485]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="385" phylum="Arthropoda" rank="species" species="coriaceus">
<emphasis id="1C97462D256A163142FEFE15FE439CD8" box="[151,391,460,484]" italics="true" pageId="1" pageNumber="385">Geodromicus coriaceus</emphasis>
<bibRefCitation id="4A72E7CE256A163143E7FE14FDA49CD9" author="Cameron, M." box="[398,608,461,485]" pageId="1" pageNumber="385" pagination="160 - 198" refId="ref3298" refString="Cameron, M. (1924) New species of Staphylinidae from India. Transactions of the Entomological Society of London, 1924, 160 - 198. https: // doi. org / 10.1111 / j. 1365 - 2311.1924. tb 03357. x" type="journal article" year="1924">Cameron, 1924: 172</bibRefCitation>
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.
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</paragraph>
<paragraph id="2E5C9A3F256A163142FEFE32FDA09F3F" blockId="1.[151,927,460,607]" box="[151,612,491,515]" pageId="1" pageNumber="385">
<treatmentCitationGroup id="0EF3BD11256A163142FEFE32FDA09F3F" box="[151,612,491,515]" pageId="1" pageNumber="385">
<treatmentCitation id="AF42BC2E256A163142FEFE32FDA49F3F" author="Cameron, M." box="[151,608,491,515]" page="144" pageId="1" pageNumber="385" year="1941">
<taxonomicName id="E9E3E1BC256A163142FEFE32FDA49F3F" authority="Cameron, 1941: 144" authorityName="Cameron" authorityPageNumber="144" authorityYear="1941" box="[151,608,491,515]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="385" phylum="Arthropoda" rank="species" species="perplexus">
<emphasis id="1C97462D256A163142FEFE32FE439F3F" box="[151,391,491,515]" italics="true" pageId="1" pageNumber="385">Geodromicus perplexus</emphasis>
<bibRefCitation id="4A72E7CE256A163143E7FE32FDA49F3F" author="Cameron, M." box="[398,608,491,515]" pageId="1" pageNumber="385" pagination="142 - 147" refId="ref3345" refString="Cameron, M. (1941) Descriptions of new Staphylinidae (Coleopt.). The Proceedings of the Royal Entomological Society of London, Series B, 10, 142 - 147. https: // doi. org / 10.1111 / j. 1365 - 3113.1941. tb 00709. x" type="journal article" year="1941">Cameron, 1941: 144</bibRefCitation>
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.
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<paragraph id="2E5C9A3F256A163142FEFDD3FD819F1E" blockId="1.[151,927,460,607]" box="[151,581,522,546]" pageId="1" pageNumber="385">
<treatmentCitationGroup id="0EF3BD11256A163142FEFDD3FD819F1E" box="[151,581,522,546]" pageId="1" pageNumber="385">
<treatmentCitation id="AF42BC2E256A163142FEFDD3FD859F1E" author="Coiffait, H." box="[151,577,522,546]" page="271" pageId="1" pageNumber="385" year="1977">
<taxonomicName id="E9E3E1BC256A163142FEFDD3FD859F1E" ID-CoL="3FPJD" authority="Coiffait, 1977: 271" authorityName="Coiffait" authorityPageNumber="271" authorityYear="1977" box="[151,577,522,546]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="385" phylum="Arthropoda" rank="species" species="palpalis">
<emphasis id="1C97462D256A163142FEFDD3FEB39F1E" box="[151,375,522,546]" italics="true" pageId="1" pageNumber="385">Geodromicus palpalis</emphasis>
<bibRefCitation id="4A72E7CE256A16314314FDD3FD859F1E" author="Coiffait, H." box="[381,577,522,546]" pageId="1" pageNumber="385" pagination="243 - 275" refId="ref3399" refString="Coiffait, H. (1977) Staphilinides recoltes au Nepal par le Professeur Franz (2 e partie). Bulletin de la Societe d'Histoire Naturelle de Toulouse, 112 (1976), 243 - 275." type="journal article" year="1977">Coiffait, 1977: 271</bibRefCitation>
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.
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<treatmentCitationGroup id="0EF3BD11256A163142FEFDF0FDAE9F7D" box="[151,618,552,577]" pageId="1" pageNumber="385">
<treatmentCitation id="AF42BC2E256A163142FEFDF0FDDA9F7D" author="Coiffait, H." box="[151,542,552,577]" page="282" pageId="1" pageNumber="385" year="1983">
<taxonomicName id="E9E3E1BC256A163142FEFDF0FDDA9F7D" ID-CoL="3NBJM" authority="Coiffait, 1983: 282" authorityName="Coiffait" authorityPageNumber="282" authorityYear="1983" box="[151,542,552,577]" class="Insecta" family="Staphylinidae" genus="Hygrogeus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="385" phylum="Arthropoda" rank="species" species="ledouxi" status="syn. nov.">
<emphasis id="1C97462D256A163142FEFDF0FE939F7C" box="[151,343,552,576]" italics="true" pageId="1" pageNumber="385">Hygrogeus ledouxi</emphasis>
<bibRefCitation id="4A72E7CE256A16314334FDF1FDDA9F7D" author="Coiffait, H." box="[349,542,552,577]" pageId="1" pageNumber="385" pagination="273 - 289" refId="ref3438" refString="Coiffait, H. (1983) Nouveaux staphylinides du Cachemire (Coleoptera). Annales de la Societe entomologique de France, New Series, 19 (3), 273 - 289. https: // doi. org / 10.1080 / 21686351.1983.12278365" type="journal article" year="1983">Coiffait, 1983: 282</bibRefCitation>
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<emphasis id="1C97462D256A1631404DFDF0FDAE9F7D" bold="true" box="[548,618,553,577]" pageId="1" pageNumber="385">
<taxonomicNameLabel id="07A4FB56256A1631404DFDF0FDAE9F7D" box="[548,618,553,577]" pageId="1" pageNumber="385">syn. n.</taxonomicNameLabel>
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<treatmentCitationGroup id="0EF3BD11256A163142FEFD9EFC5B9F63" box="[151,927,583,607]" pageId="1" pageNumber="385">
<taxonomicName id="E9E3E1BC256A163142FEFD9EFE599F63" ID-CoL="3NBHT" baseAuthorityName="Cameron" baseAuthorityYear="1924" box="[151,413,583,607]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="385" phylum="Arthropoda" rank="species" species="coriaceus">
<emphasis id="1C97462D256A163142FEFD9EFE599F63" box="[151,413,583,607]" italics="true" pageId="1" pageNumber="385">Hygrodromicus coriaceus</emphasis>
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: 
<treatmentCitation id="AF42BC2E256A163143C2FD9EFD9B9F63" author="Zerche, L." box="[427,607,583,607]" page="120" pageId="1" pageNumber="385" year="1992">
<bibRefCitation id="4A72E7CE256A163143C2FD9EFD9B9F63" author="Zerche, L." box="[427,607,583,607]" pageId="1" pageNumber="385" pagination="105 - 142" refId="ref3633" refString="Zerche, L. (1992) Zur Taxonomie, Phylogenie und Verbreitung der Hygrogeus - Gruppe Mittel- und Zentralasiens (Coleoptera: Staphylinidae: Omaliinae). Annalen des Naturhistorischen Museums in Wien, 93 (B), 105 - 142." type="journal article" year="1992">Zerche 1992: 120</bibRefCitation>
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, 
<treatmentCitation id="AF42BC2E256A16314003FD9EFCE39F63" author="Shavrin, A. V." box="[618,807,583,607]" page="338" pageId="1" pageNumber="385" year="2021">
<bibRefCitation id="4A72E7CE256A16314003FD9EFCE39F63" author="Shavrin, A. V." box="[618,807,583,607]" pageId="1" pageNumber="385" pagination="335 - 348" refId="ref3483" refString="Shavrin, A. V. (2021) New taxonomic and faunistic data of the genus Hygrodromicus Tronquet, 1981 of the Himalayan Region (Coleoptera: Staphylinidaae: Omaliinae: Anthophagini). In: Hartmann, M., Barclay, M. &amp; Weipert, J. (Eds.), Biodiversitat und Naturausstattung im Himalaya VII. Verein der Freunde und Forderer des Naturkundmuseums Erfurt e. V., Erfurt, pp. 335 - 348." type="book chapter" year="2021">Shavrin 2021: 338</bibRefCitation>
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, 
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<bibRefCitation id="4A72E7CE256A1631415BFD9EFC5F9F63" author="Shavrin, A. V." box="[818,923,583,607]" pageId="1" pageNumber="385" pagination="236 - 246" refId="ref3567" refString="Shavrin, A. V. (2024) New species and records of Hygrodromicus Tronquet, 1981 and Paratrichodromeus Zerche, 1992 from the Middle Asia, the Himalayan Region and China (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa, 5446 (2), 236 - 246. https: // doi. org / 10.11646 / zootaxa. 5446.2.5" type="journal article" year="2024">2024: 237</bibRefCitation>
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.
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</paragraph>
</subSubSection>
<subSubSection id="66F9C9B4256A163142FEFD54FD669E2F" pageId="1" pageNumber="385" type="discussion">
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<emphasis id="1C97462D256A163142FEFD54FECF9F9B" bold="true" box="[151,267,653,679]" pageId="1" pageNumber="385">Remarks.</emphasis>
<taxonomicName id="E9E3E1BC256A1631437AFD57FE239F9B" authorityName="Coiffait" authorityYear="1983" box="[275,487,653,679]" class="Insecta" family="Staphylinidae" genus="Hygrogeus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="385" phylum="Arthropoda" rank="species" species="ledouxi">
<emphasis id="1C97462D256A1631437AFD57FE239F9B" box="[275,487,653,679]" italics="true" pageId="1" pageNumber="385">Hygrogeus ledouxi</emphasis>
</taxonomicName>
based on the single 
<typeStatus id="F158249D256A163140A6FD54FCF59F9B" box="[719,817,653,679]" pageId="1" pageNumber="385" type="holotype">holotype</typeStatus>
was originally described from “Cachemire, Pahalgam, Sheshnag…”. Based on the original description and the shape of the aedeagus (
<figureCitation id="B6D886BA256A16314679FD68FB749FF7" box="[1040,1200,689,715]" captionStart="FIGURES 1–2" captionStartId="2.[151,264,1207,1231]" captionTargetBox="[157,1430,281,1175]" captionTargetId="figure-60@2.[151,1436,270,1181]" captionTargetPageId="2" captionText="FIGURES 1–2. Habitus of Hygrodromicus: 1–H. incrassatus (paratype), 2–H. splendidus (Kazakhstan, Ketmen` Mts.). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12758946" httpUri="https://zenodo.org/record/12758946/files/figure.png" pageId="1" pageNumber="385">Figs 2I and 2J</figureCitation>
in 
<bibRefCitation id="4A72E7CE256A163146BDFD68FAA99FF7" author="Coiffait, H." box="[1236,1389,689,715]" pageId="1" pageNumber="385" pagination="273 - 289" refId="ref3438" refString="Coiffait, H. (1983) Nouveaux staphylinides du Cachemire (Coleoptera). Annales de la Societe entomologique de France, New Series, 19 (3), 273 - 289. https: // doi. org / 10.1080 / 21686351.1983.12278365" type="journal article" year="1983">Coiffait 1983</bibRefCitation>
), 
<taxonomicName id="E9E3E1BC256A163147E8FD6BFF2D9FD3" authorityName="Coiffait" authorityYear="1983" class="Insecta" family="Staphylinidae" genus="Hygrogeus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="385" phylum="Arthropoda" rank="species" species="ledouxi">
<emphasis id="1C97462D256A163147E8FD6BFF2D9FD3" italics="true" pageId="1" pageNumber="385">H. ledouxi</emphasis>
</taxonomicName>
is conspecific with the Himalayan 
<taxonomicName id="E9E3E1BC256A1631401CFD0FFCC09FD3" baseAuthorityName="Cameron" baseAuthorityYear="1924" box="[629,772,726,751]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="385" phylum="Arthropoda" rank="species" species="coriaceus">
<emphasis id="1C97462D256A1631401CFD0FFCC09FD3" box="[629,772,726,751]" italics="true" pageId="1" pageNumber="385">H. coriaceus</emphasis>
</taxonomicName>
, also known from Kashmir (
<bibRefCitation id="4A72E7CE256A1631462FFD0CFB259FD3" author="Shavrin, A. V." box="[1094,1249,725,751]" pageId="1" pageNumber="385" pagination="335 - 348" refId="ref3483" refString="Shavrin, A. V. (2021) New taxonomic and faunistic data of the genus Hygrodromicus Tronquet, 1981 of the Himalayan Region (Coleoptera: Staphylinidaae: Omaliinae: Anthophagini). In: Hartmann, M., Barclay, M. &amp; Weipert, J. (Eds.), Biodiversitat und Naturausstattung im Himalaya VII. Verein der Freunde und Forderer des Naturkundmuseums Erfurt e. V., Erfurt, pp. 335 - 348." type="book chapter" year="2021">Shavrin 2021</bibRefCitation>
, 
<bibRefCitation id="4A72E7CE256A16314685FD0CFAED9FD3" author="Shavrin, A. V." box="[1260,1321,725,751]" pageId="1" pageNumber="385" pagination="236 - 246" refId="ref3567" refString="Shavrin, A. V. (2024) New species and records of Hygrodromicus Tronquet, 1981 and Paratrichodromeus Zerche, 1992 from the Middle Asia, the Himalayan Region and China (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa, 5446 (2), 236 - 246. https: // doi. org / 10.11646 / zootaxa. 5446.2.5" type="journal article" year="2024">2024</bibRefCitation>
). Thus, 
<taxonomicName id="E9E3E1BC256A163147E8FD0FFF2D9E2F" authorityName="Coiffait" authorityYear="1983" class="Insecta" family="Staphylinidae" genus="Hygrogeus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="385" phylum="Arthropoda" rank="species" species="ledouxi">
<emphasis id="1C97462D256A163147E8FD0FFF2D9E2F" italics="true" pageId="1" pageNumber="385">H. ledouxi</emphasis>
</taxonomicName>
is synonymized with the latter species.
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