New species of Nymphites Haase (Neuroptera: Nymphidae) from the Middle Jurassic of China, with a redescription of the type species of the genus
Author
Shi, Chaofan
Author
Makarkin, Vladimir N.
Author
Yang, Qiang
Author
Archibald, S. Bruce
Author
Ren, Dong
text
Zootaxa
2013
3700
3
393
410
journal article
10.11646/zootaxa.3700.3.4
749f4c56-ff8e-496a-9b83-8befda755dbe
1175-5326
248765
E903900B-9509-4602-BEF2-483280945E73
Nymphites priscus
(Weyenbergh, 1869)
(
Figs 1
,
2
)
Insecte: Winkler 1865: 429 (listed); Winkler 1896: 314 (listed).
Hemerobius priscus
Weyenbergh 1869a: 264
; Pl. 34, Figs 13, 14 (original description); Weyenbergh 1869b: 236 (listed); Weyenbergh 1874: 97 (listed); Oppenheim 1888: 226, 227 (as a synonym of
Chrysopa excelsa
Hagen, 1862
[=
Osmylites excelsa
Oppenheim, 1888
]); Scudder 1891: 148 (listed); Martins-Neto & Vulcano 1989: 369, 370 (as a synonym of
Nymphites braueri
Haase, 1890
).
Névroptère: Meunier 1897: 238 (notes).
Nymphites priscus
: Haase 1890: 23
(taxonomic notes); Handlirsch
1906–1908
[1906]: 609 (taxonomic notes); Martynova 1949: 167 (listed); Lambkin 1988: 450, 453 (mentioned); Martins-Neto 1992: 120 (mentioned); Makarkin & Archibald 2003: 175 (taxonomic notes); Ponomarenko 2003: 90 (mentioned); Makarkin & Menon 2005: 810 (taxonomic notes).
FIGURE 1.
Nymphites priscus
(Weyenbergh, 1869)
, holotype. A, part. B, counterpart. Scale bar = 5 mm.
FIGURE 2.
Nymphites priscus
(Weyenbergh, 1869)
, holotype. A, right hind wing photograph. B, drawing of hind wing venation. Dotted line indicates both ScP and RA, which are not distinctly separate here by blurred preservation. Scale bar = 5 mm.
Diagnosis.
Distinguished from
Nymphites bimaculatus
sp. nov.
and from
Nymphites
sp. A by the absence of crossveins between distal branches of CuA and branches of MP.
Description.
Thorax fragmentarily preserved; details not discernible.
Hind
wing approximately
29 mm
long as preserved (estimated complete length
31 mm
); 8.0 mm wide as preserved (estimated complete width
8.5 mm
). Margins poorly-preserved; trichosors not visible. Costal space narrow. Subcostal veinlets simple in proximal 2/3, some distal veinlets apparently shallowly forked. ScP, RA preserved as very closely approaching veins; subcostal space not visible. Preserved veinlets of ScP + RA dichotomously branched.
RP
smooth, only slightly zigzagged; originating near wing base. Preserved crossveins between RA,
RP
before fusion of ScP + RA widely spaced, rather regularly arranged.
RP
with 11 or 12 branches (poorly preserved); at least
RP
1,
RP
2 deeply forked. Crossveins in radial space relatively scarce, irregularly spaced proximally; rare or absent distally. Origin of M not preserved. MA dichotomously branched relatively close to wing margin. MP pectinately branched, with four branches, two deeply forked; one preserved crossvein evident between them. CuA long, pectinately branched with 11 long branches, at least one deeply forked; four-six crossveins between proximal branches forming gradate series. CuP incompletely preserved, short, apparently deeply forked. Anal veins not preserved. Crossveins between
RP
1 and MA, MA and MP, MP and CuA rather scarce, irregularly spaced.
Type
material
.
Holotype
No. 15453 (part; formerly No. 10336), No. 15454 (counterpart), deposited in TM. A poorly-preserved specimen with two incomplete hind wings.
Type
locality and horizon.
Solnhofen,
Germany
; Late Jurassic, early Tithonian (Solnhofen Formation).
Discussion.
Handlirsch (
1906–1908
) thought that these are forewings, apparently following Weyenbergh (1869a) and Haase (1890).