Two new species of Lysianassidae Dana, 1849 from Australia: Riwo zeidleri and Socarnella delectabilis (Crustacea: Peracarida: Amphipoda)
Author
Hughes, L. E.
Author
Lowry, J. K.
text
Zootaxa
2015
3936
1
journal volume
10.11646/zootaxa.3936.1.4
f4db1ead-5e93-45cd-af4a-45f5c8d2c2ca
1175-5326
244435
0E9B41A5-B557-4CBF-943A-EC082666B79F
Riwo zeidleri
sp. nov.
(
Figs 1
,
2
)
Type
material
.
Holotype
, female,
4.1 mm
, dissected, 3 slides,
SAMA
C
8391, 200 m
offshore, north point of Marum Island, Sir Joseph Banks Group, South
Australia
(
34°31’S
136°15’E
),
6 m
, on sponge and under rocky reef,
11 January 1984
, coll. W. Zeidler (SAM-14);
paratypes
5 specimens
(inc. male
2.4 mm
),
SAMA
C8392, same location as
holotype
.
Type
locality
.
200 m
offshore, north point of Marum Island, Sir Joseph Banks Group, South
Australia
(
34°31’S
136°15’E
).
Etymology
. Named for Wolfgang Zeidler, the collector of the species, in appreciation for his elucidating studies of hyperiidean amphipods.
Description
.
Holotype
female,
4.1 mm
, SAMA C8391;
paratype
male,
2.4 mm
, SAMA C8392. Colour whitish with small black chromatophores covering pereon, coxae and epimera, but concentrated anteriorly on pereon segments to give a striped effect; body without setae.
Head
as deep as long; lateral cephalic lobe large, broadly rounded; rostrum small; eyes subovate, black.
Antenna 1
medium length, 0.3 × body; peduncular article 1 short, length 1.1 × width; article 2 short, 0.25 × article 1; article 3 short, 0.2 × article 1; accessory flagellum long, 0.4 × primary flagellum,
5-articulate
,
article 1 subequal to article 2 length
; flagellum 9-articulate, callynophore weak 1-field, without posterodistal slender or robust setae, without flagellar robust setae, calceoli absent.
Antenna 2
subequal in length to antenna 1; peduncle with brush setae present, strongly geniculate between peduncular articles 3–4; article 3 long, 0.8 × article 4; peduncular articles 4–5 not enlarged; flagellum 8-articulate, calceoli absent.
Mouthpart bundle
: subquadrate.
Labrum
epistome concave, separate from upper lip; upper lip produced, rounded.
Mandible
incisors symmetrical, large, with convex margins; lacinia mobilis absent; accessory setal row without distal setal tuft, left and right rows each with 3 long, slender, multiserrate robust setae; molar developed as small, smooth setose flap; mandibular palp attached proximally, article 1 short, length 1.1 × width; article 2 elongate, slender, length 4.3 × width, 1.8 × article 3, with 5 posterodistal A2-setae; article 3 slender, blade-like, long, length 3 × width, without A3- or B3-setae, with 1 proximal and 2 distal D3-setae on posterior margin, and 2 apical E3-setae.
Maxilla 1
inner plate narrow with 1 simple apical seta (broken); outer plate with 10 setal-teeth in modified 6/5 arrangement; palp large, 2-articulate,
with 7 short terminal conate setae and 1 subterminal seta
,
distomedial margin not serrate
.
Maxilla 2
inner and outer plates broad, subequal in length.
Maxilliped
inner plate large, subrectangular, with 3 apical nodular setae, oblique setal row reduced with 2 setae; outer plate small, subrectangular, without apical slender or robust setae, 8 vestigial medial robust setae; palp large, 4-articulate; article 2 broad, length 2.9 × width, 1.1 × article 3; article 3 long, slender, length 2 × width; dactylus well-developed, with 1 subterminal seta, unguis present.
Pereon.
Oostegites from gnathopod 2 to pereopod 5. Gills from gnathopod 2 to pereopod 7, not pleated.
Gnathopod 1
simple; coxa large, 0.9 × coxa 2, anterior margin straight, anteroventral corner weakly produced, subquadrate, posterior margin straight;
basis
long, slender,
length 2.6 × width
, anterior margin smooth, without setae; ischium short, length 1.1 × width; merus, posterior margin with patch of short setae and a few simple slender setae; carpus subrectangular, short, length 1.9 × width, subequal in length to propodus, without denticulate patch near posterodistal margin; propodus small, subrectangular, length 2.1 × width, tapering distally, posterior margin smooth, straight, with few slender setae, without denticulate patch near posterior margin, palm absent; dactylus simple,
without subterminal spine
.
Gnathopod 2
minutely subchelate; coxa large, subequal in size to coxa 3; ischium long,
length 3.1 × width
; carpus long,
length 2.4 × width
, posterior margin straight; propodus subrectangular, short, length 1.6 × width, palm obtuse, with straight, serrate margin, posterodistal corner without robust setae;
dactylus
reaching corner of palm,
posterior margin smooth
.
Pereopod 3
coxa large; merus weakly expanded anteriorly; merus and carpus posterior margin lined with slender setae; propodus with 3 long slender setae and 2 distal locking setae along posterior margin; dactylus recurved.
Pereopod 4
coxa deeper than wide, with large posteroventral lobe, anterior margin rounded, posterior margin sloping anteriorly; merus weakly expanded anteriorly; merus and carpus posterior margin lined with slender setae; propodus with 1 slender seta and 1 distal locking seta along posterior margin; dactylus recurved.
Pereopod
5 coxa equilobate; basis expanded with posterior margin minutely crenate; merus distally expanded, margin sloping proximally, straight distally; propodus with 2 distal locking setae along anterior margin; dactylus recurved.
Pereopod 6
coxa small, not lobate posteriorly; basis expanded posteriorly with minutely crenate posterior margin, without anteroventral lobe; merus expanded posteriorly margin sloping proximally, straight distally; propodus with 3 small robust setae and 2 distal locking setae along anterior margin; dactylus recurved.
Pereopod 7
basis expanded posteriorly, posterior margin slightly rounded, minutely crenate, posteroventral corner rounded, posteroventral margin rounded; merus not expanded posteriorly; propodus with 3 small robust setae and 2 distal locking setae along anterior margin; dactylus recurved.
FIGURE 1
.
Riwo zeidleri
sp. nov.
, holotype female, 4.1 mm, SAMA C8391, Marum Island, Sir Joseph Banks Group, South Australia. Scale 0.1 mm.
FIGURE 2
.
Riwo zeidleri
sp. nov.
, holotype female, 4.1 mm, SAMA C8391, Marum Island, Sir Joseph Banks Group, South Australia. Scale 0.2 mm.
Pleon.
Pleonites 1–3
dorsally smooth.
Epimeron 1
posteroventral corner broadly rounded.
Epimeron 2
posteroventral corner subquadrate.
Epimeron 3
subquadrate, posteroventral corner rounded.
Urosomites 1–3
dorsally smooth.
Uropod 1
without long fine setae; peduncle with 2 dorsolateral and 2 apicomedial robust setae; outer ramus subequal to inner ramus length; outer ramus with 1 lateral robust seta; inner ramus without robust setae.
Uropod 2
without fine setae; peduncle without dorsolateral setae, with 2 apicomedial robust setae; rami subequal in length,
outer ramus without dorsal robust setae;
inner ramus without robust setae, without constriction.
Uropod 3
peduncle long, length 2 × width, without dorsolateral flange or robust setae; rami lanceolate, inner ramus reduced, 0.6 × outer ramus; outer ramus 2-articulate, article 2 short, 2.1 × width; rami without robust setae.
Telson
length 1.1 × width,
weak emarginated notch,
without dorsal robust setae, with 2 marginal penicillate seta.
Remarks
.
Riwo zeidleri
sp. nov.
, the second species in the genus, can be separated from
Riwo mizeui
Lowry & Stoddart, 1995
from
Papua New Guinea
, by the antenna 1 accessory flagellum with 5 articles (4-articulate in
R. mizeui
), maxilla 1 palp article two with 7 connate robust setae (only
3 in
R. mizeui
) and by the weakly emarginated telson (notched in
R. mizeui
).
Distribution.
Marum Island, Sir Joseph Banks Group, South
Australia
(current study).