A New Fossil Sucinorhagonycha (Coleoptera: Cantharidae) From Baltic Amber With Visible Female Gonostyli
Author
Fanti, Fabrizio
text
Fossil Imprint
2023
2023-12-22
79
2
152
155
http://dx.doi.org/10.37520/fi.2023.009
journal article
10.37520/fi.2023.009
2533-4069
10481005
Sucinorhagonycha carsteni
fANTI
sp. nov.
Text-fig. 1
H o l o t y p e. Female, inclusion in Baltic amber, housed at the GPIH (
Germany
) with the identification number GPIH no. 5090, CCGG no. 8696 (ex Jonas Damzen Collection JDC12143),
Text-fig. 1.
E t y m o l o g y. Species named in honor of Carsten
Gröhn, a well-known German amber collector.
T
y p e l o c a l i t y.
Amber
mine near
Yantarny
settlement,
Sambian Peninsula
,
Kaliningrad region
,
Russia
.
T y p e h o r i z o n. Sediments containing Baltic amber.
A g e. Middle Eocene (Lutetian) (47.8–41.2 Ma) to late
Eocene (Priabonian) (37.8–33.9 Ma).
D i a g n o s i s. Soft bodied soldier beetle with prognathous head, last maxillary palpomere securiform, 12-segmented antennae that are filiform without modified antennomeres and without rami, pronotum elongated without lobes on the sides, long smooth elytra without costae, tarsal formula 5-5-5, and simple claws without lobes or denticles.
C o m p a r i s o n.
Sucinorhagonycha maryae
M.G.
PANKOWSKI
et
FANTI
, 2023
has a flat and transverse pronotum (
Pankowski and Fanti 2023
); the pronotum of
S. samsockorum
FANTI
et M.K.
PANKOWSKI
, 2018
has a clearly raised area in the center (
Fanti and Pankowski 2018
);
S. groehni
KAZANTSEV
, 2020
has a pronotum with somewhat produced anterior margin and prominent posterior bulges (
Kazantsev 2020
); and in
S. kulickae
KUŚKA
, 1996
the pronotum has a conspicuous central linear depression extending from anterior to posterior margin and distinct protuberances on both sides (
Kuśka 1996
,
Kubisz 2000
).
S. fabrizioi
M.G.
PANKOWSKI
, 2023
presents a pronotum with a clearly bulged area near the middle (Pankowski 2023). The new species
S. carsteni
is therefore well differentiated because it does not have any of these combinations of characters.
D e s c r i p t i o n. Adult, winged, elongated. Female, defined on the basis of its rounded last sternite and visible ovipositor (gonostyli). Body length
5.7 mm
: abdomen (elytra)
3.9 mm
; thorax
1.1 mm
; head
0.7 mm
. Antenna
3.4 mm
. Body entirely dark brown.
Head slightly elongated, prognathous, not completely exposed, transverse, with shallow punctation and short setae. Eyes rounded, small, convex, inserted in the lateral part of the head. Mandibles robust, elongated, falciform. Maxillary palps 4-segmented and segments unequal in length; last palpomere securiform, elongated, rounded apically. Labial palps 3-segmented; last palpomere elongated, securiform. Antennae 12-segmented, filiform, inserted dorsally and between eyes, relatively short, extending beyond half of the elytra, all segments pubescent; antennomere I (scape) robust, elongated, slightly club-shaped; antennomere II robust, short; antennomere III robust, sturdier and about 1.4 times longer than the second; antennomere IV slightly thinner and slightly longer than antennomere III; antennomeres V–X subequal, slightly shorter than previous one, enlarged at external margin; antennomere XI elongated-rounded; antennomere XII elongated, rounded at apex. Pronotum longer than wide, very slightly wider than the head, corners rounded, posterior margin almost straight and with prominent border, anterior margin rounded and with delicate border, surface rather flat and with very small bulged areas in the posterior part, and with a transverse depression following the bulged areas and reaching the posterior margin, surface with punctation and strongly pubescent. Scutellar shield triangular-shaped, wide basally with rounded apex. Elytra very elongated, slender, parallel-sided, completely covering the last abdominal segments, wider than pronotum, apex rounded, surface equipped with long setae and slightly wrinkled. Posterior wings completely covered by elytra. Metasternum elongated, posteriorly slightly rounded; sternites and tergites wide, narrow, slightly punctate, pubescent; last sternite wide, narrow and strongly transverse, with apical margin undulate, equipped with two short central lobes, very robust and obtuse apically; ovipositor with two visible very thin, long and well-developed gonostyli. Legs short, massive, strongly pubescent; coxae massive; trochanters elongated; femora cylindrical, enlarged and slightly curved; tibiae shorter than femora, cylindrical, thin. Tarsi 5-segmented and with setae; first tarsomere elongated and very robust, about 2.0 times longer than second; second tarsomere short and robust; third tarsomere shorter than second; fourth tarsomere strongly bilobed at sides with the lobes very long and obtuse apically; fifth tarsomere very elongated, thin, strongly curved; claws simple, long, curved, and without lobes or denticles. Male unknown.
Text-fig. 1.
Sucinorhagonycha carsteni
FANTI
sp. nov.
, Baltic amber, holotype (GPIH no. 5090, CCGG no. 8696). a: Dorsal view, scale bar 1.0 mm. b: Ventral view, scale bar 1.0 mm. c: Detail of antennae, scale bar 0.5 mm. d: Detail of last sternites with gonostyli, scale bar 0.3 mm (© Jonas & Aleksej Damzen).
R e m a r k s. The yellow amber piece measures approximately 41 × 20 ×
8 mm
. The inclusion is complete and clearly visible. It has white emulsion only on the ventral side.
S y n i n c l u s i o n s. Air bubbles, plant remains, stellate hairs,
Trichoneura
sp.
(
Diptera
Limoniidae
).