Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java Author Nurinsiyah, Ayu Savitri 5AFBFEFB-B243-49CC-AACC-248B2F007BC8 Museum Zoologicum Bogoriense, Research Center for Biology, Indonesian Institute of Science, Cibinong, West Java, Indonesia. ayu_nurinsiyah@yahoo.com & ayus002@lipi.go.id Author Neiber, Marco T. 8F6006F5-D880-4841-AC32-B97E18D7BB5C Zoological Museum, Center of Natural History, University of Hamburg, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany mneiber@hotmail.de Author Hausdorf, Bernhard FEDE92C0-108D-4315-8F86-9F2CFF1CB906 Zoological Museum, Center of Natural History, University of Hamburg, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany. hausdorf@zoologie.uni-hamburg.de text European Journal of Taxonomy 2019 2019-05-21 526 1 73 journal article 10.5852/ejt.2019.526 2da65c0a-9c48-43f5-ada3-e088de28dec1 2118-9773 3152217 3FFC527D-E9CA-4D9F-BF59-076FA60171AC Landouria ciliocincta ( Möllendorff, 1897 ) Figs 45 , 54 , 69 , 83–84 ; Tables 3–4 Plectotropis ciliocincta Möllendorff, 1897: 67 (“ Java ”; more exactly ( Zilch 1966 ): “O-Java: Zuider- Gebirge”). Landouria ciliocincta – van Benthem Jutting 1950: 466 (in part). — Zilch 1966: 293 , pl. 7, fig. 1 (in part). — Dharma 2015: 8 , figs 2, 4 (in part). — Nurinsiyah et al. 2016: 559 . Diagnosis Landouria ciliocincta is characterized by a whitish, almost disc-like, keeled shell with an only slightly elevated spire, a reddish-brown band at the angular periphery and a wide umbilicus, a long, cylindrical penis with a small bulge at its proximal end, and a short, very broad flagellum, which tapers towards its proximal end. Material examined Lectotype INDONESIA • lectotype (designated by Zilch 1966 ); East Java , Zuider (Southern) Mountains ; SMF 9176 a. Paralectotype INDONESIA • 1 spec. ; same data as for lectotype; SMF 9716 b. Other material INDONESIA – East Java • 1 spec. (voucher for Dharma 2015 ); Malang , Ngliyep ; 8°23′ S , 112°25′ E ; near sea level ; BDJ • 1 spec. (det. anat.) (voucher for Nurinsiyah et al. 2016 ); Malang , Kondangmerak , nature/protected forest ; 8°23′22″ S , 112°31′02″ E ; 51 m a.s.l.; MZB 19211 • 1 spec. (voucher for Nurinsiyah et al. 2016 ); Malang , Kondangmerak , nature/protected forest ; 8°23′21″ S , 112°31′05″ E ; 100 m a.s.l.; MZB 17645 • 2 spec. ; Tengger Mountains ; ZMB 45408 . Description SHELL (Figs 83–84; Tables 3–4 ). Almost disc-like with slightly elevated spire, with 5–6 moderately convex whorls; protoconch almost smooth; teleoconch with fine, irregular wrinkles, indistinct incised spiral lines and sparse scaly processes, mainly at periphery; tuberculate around umbilicus; whitishcorneous with a brownish band at periphery; body whorl keeled; aperture rounded rhombic; upper insertion of peristome descending; peristome expanded, reflexed and thickened; umbilicus slightly eccentric, wide, comprising 22–30% of shell diameter, hardly obscured by columellar edge. GENITALIA ( Figs 45 , 69 ; Table 4 ). Atrium short; penis long, cylindrical, with a small bulge at its proximal end, without penis coecum; membranaceous sheath around distal part of penis connected with flagellum by tissue; penial retractor runs from diaphragm to middle part of cylindrical epiphallus; flagellum short, very broad, without node, tapering towards its proximal end; vas deferens narrow and long; vagina long; oviduct short, broad, proximally tapering; peduncle of bursa copulatrix subdivided into a broad, proximally tapering distal part and a narrower, longer proximal part, ending in an elongate oval bursa beside albumen gland. Right ommatophoral retractor runs between penis and vagina. Remarks Landouria ciliocincta shares with L. pakidulan sp. nov. and L. menorehensis sp. nov. the whitish shell with a brown band at the periphery. For differences, see remarks under L. pakidulan sp. nov. Distribution Landouria ciliocincta is known from Malang Regency in East Java (Fig. 54). Records from other areas of Java ( Oostingh 1923 ; van Benthem Jutting 1941 , 1950 ; Zilch 1966 ; Dharma 2015 ) and from Sumatra ( van Benthem Jutting 1959 ; Heryanto 2013 ) probably refer to other species.