Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java Author Nurinsiyah, Ayu Savitri 5AFBFEFB-B243-49CC-AACC-248B2F007BC8 Museum Zoologicum Bogoriense, Research Center for Biology, Indonesian Institute of Science, Cibinong, West Java, Indonesia. ayu_nurinsiyah@yahoo.com & ayus002@lipi.go.id Author Neiber, Marco T. 8F6006F5-D880-4841-AC32-B97E18D7BB5C Zoological Museum, Center of Natural History, University of Hamburg, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany mneiber@hotmail.de Author Hausdorf, Bernhard FEDE92C0-108D-4315-8F86-9F2CFF1CB906 Zoological Museum, Center of Natural History, University of Hamburg, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany. hausdorf@zoologie.uni-hamburg.de text European Journal of Taxonomy 2019 2019-05-21 526 1 73 journal article 10.5852/ejt.2019.526 2da65c0a-9c48-43f5-ada3-e088de28dec1 2118-9773 3152217 3FFC527D-E9CA-4D9F-BF59-076FA60171AC Landouria tholiformis sp. nov. urn:lsid:zoobank.org:act: C083E997-420A-4CAA-8A67-088C02177B83 Figs 7 , 20 , 34 , 51–52 ; Tables 3–4 Diagnosis Landouria tholiformis sp. nov. is characterized by a brownish-corneous, sharply keeled shell with scaly processes all over the shell, with a bulge at the proximal end of the penis forming a short coecum, and a crenated flagellum, which tapers abruptly at its proximal end. Etymology Landouria tholiformis sp. nov. is named after its dome-shaped shell (Latin: ʻtholusʼ = ʻdomeʼ, ʻformaʼ = ʻshapeʼ; used as an adjective). Material examined Holotype INDONESIA • holotype (det. anat.); Gunung Kidul , Tepus , near Poktunggal Beach , agroforestry ; 8°08′42″ S , 110°37′52″ E ; 137 m a.s.l. ; 9 Aug. 2014 ; A.S. Nurinsiyah , F.L.H. Irsyad , F.J. Sari and E. Nurlaela leg.; D = 8.7 mm , H = 5.2 mm ; MZB 17999 . Paratypes INDONESIA • 8 spec. ; Yogyakarta , Gunung Kidul , Tepus , Pego Hill , agroforestry ; 8°06′10″ S , 110°37′50″ E ; 230 m a.s.l. ; MZB 19299 . Description SHELL ( Figs 7 , 51 ; Tables 3–4 ). Depressed conical, with a dome-shaped spire, with 6–6.25 slightly convex whorls; protoconch almost smooth; teleoconch with fine, irregular wrinkles; apical side with indistinct incised spiral lines, umbilical side with more distinct spiral lines; scaly processes all over shell; tuberculate around umbilicus; brownish-corneous; body whorl sharply keeled; aperture rounded rhombic; upper insertion of peristome descending; peristome expanded, reflexed and thickened; umbilicus slightly eccentric, wide, comprising 32–40% of shell diameter, hardly obscured by columellar edge. GENITALIA ( Figs 20 , 34 ; Table 4 ). Atrium short; penis long, becoming slightly broader proximally, with a bulge at proximal end forming a short coecum; membranaceous sheath around distal part of penis connected with epiphallus by tissue; penial retractor runs from diaphragm to proximal part of cylindrical epiphallus; flagellum moderately long, broadest in its middle part, crenated, without node, abruptly tapering at its proximal end; vas deferens narrow and long; vagina moderately long; oviduct short, broad, proximally tapering; peduncle of bursa copulatrix subdivided into a broader, almost cylindrical distal part that abruptly passes into a narrower and longer proximal part that ends in an elongate oval bursa beside albumen gland. Right ommatophoral retractor runs between penis and vagina. Remarks Landouria tholiformis sp. nov. clusters in the phylogeny ( Fig. 1 ) with L. abdidalem sp. nov. (Fig. 56 ). It differs from that species in the more depressed (D/H 1.77–1.88 vs 1.93–2.14 in L. abdidalem sp. nov. ), less sharply keeled shell with scaly processes all over the teleoconch ( Fig. 7 ), in the flagellum, which is slightly longer than the short epiphallus, and in having a bulge at the proximal end of the penis (perhaps this is an individual abnormity). Landouria tholiformis sp. nov. occurs sympatrically with L. sewuensis sp. nov. (Fig. 58), from which it differs in the less sharply keeled shell with a dome-shaped spire, a more distinct granular microsculpture, with scaly processes all over the shell ( Fig. 7 ) and less distinct spiral lines at the umbilical side, the bulge at the proximal end of the penis that forms a short coecum, and a narrow, crenated flagellum. The flagellum of L. sewuensis sp. nov. is broader than that of L. tholiformis sp. nov. and equipped with two rows of tubercles. There is a globular bulge in the distal part of the vagina in L. sewuensis sp. nov. which is absent in L. tholiformis sp. nov. Distribution Landouria tholiformis sp. nov. is endemic to the Gunung Sewu karst region in Gunung Kidul Regency (Fig. 52).