Taxonomic revision of the spider genera Agyneta and Tennesseellum (Araneae, Linyphiidae) of North America north of Mexico with a study of the embolic division within Micronetinae sensu Saaristo & Tanasevitch 1996 Author Dupérré, Nadine text Zootaxa 2013 3674 1 1 189 journal article 10.11646/zootaxa.3674.1.1 9c8e7b58-7cc0-407f-a8e9-d76311dcd290 1175-5326 283954 981F80ED-96D7-40C7-8A3C-677954416A2E Agyneta vinki new species Figs 213–222 , map 14 Type material: Male holotype from Oregon, Imber, 24.ix.1964 , W. Ivie ( AMNH ). EXAMINED . Etymology: The specific name is a patronym in honor of Dr. Cor Vink , Kiwi arachnologist and wonderful friend. Diagnosis: Males are diagnosed from most species in the genus by the three prongs of the lamella characteristica ( Fig. 213 ). From A. panthera by the absence of cymbial tubercles ( Fig. 214 ), elongated in the latter ( Fig. 224 ). Females are diagnosed from all species by their wide proximal part of scape constricted medially ( Fig. 219 ). From A. panthera by their shorter median part of scape (Fig, 221), much longer in the latter ( Fig. 231 ). Description: Male : Total length 1.74; carapace length 0.88, width 0.71. CEPHALOTHORAX: Carapace dark orange-brown, shiny, finely reticulate; suffused with dark gray along radiating lines, margin; trident mark present. Sternum dark brown, suffused with dark gray. Clypeus height 2. Chelicerae orange with transverse dark gray suffused band, excavated; ~17 seta-tipped tubercles; promargin three teeth, retromargin three denticles, both margins with triangular, elongated projection near base of fang. Cheliceral stridulatory organ ~35 striae, well spaced apically, narrowing basally. ABDOMEN: Patterned, vertical gray band reaching halfway on the abdomen and five transverse gray chevrons ( Fig. 217 ). LEGS: Orange; leg I total length: 4.38; leg III total length: 2.72; Tm I: 0.24, Tm IV: absent. GENITALIA: Palpal retrolateral tibial apophysis small, pointed; dorsal tibial apophysis reduced, rugose; two retrolateral trichobothria and one dorsal ( Fig. 213 ). Cymbium triangular; glabrous depression present ( Fig. 213 ); cymbial tubercles absent; prolateral notch shallow ( Fig. 214 ). Paracymbium apical pocket long, anterior pocket short and curved, making a medium cover, posterior pocket long ( Fig. 213 ). Embolus tip pointed, with serrated extension; basally with one large prong; ventrally with numerous small spines; Fickert’s gland absent; ventral lamella small, pointed; thumb going slightly over the embolus proper ( Fig. 215 ). Embolus proper set apically, dorsal part larger, not serrated ( Fig. 215 ). Anterior terminal apophysis with pointed tip and with numerous long protrusions; posterior terminal apophysis small with blunt tip; lamella characteristica large, with two pointed prongs and a blunt one ( Fig. 216 ). Female : Total length 2.14; carapace length 0.85, width 0.65. MAP. 14. Localities of Agyneta vinki n. sp. CEPHALOTHORAX: Same coloration as male. Chelicerae orange; promargin 5 teeth, retromargin 5 denticles. Cheliceral stridulatory organ ~56 striae, well spaced apically, narrowing basally. ABDOMEN: Patterned, with short vertical gray band and five broken chevons ( Fig. 218 ). LEGS: Same as male; palp tibia, metatarsus suffused with gray, tarsus normal, claw absent; leg I total length: 3.46; leg III total length: 2.39; Tm I: 0.25, Tm IV: absent. GENITALIA: Epigynum with wide proximal part of scape, constricting and then enlarging; epigynal slits rounded; pit hook depression shallow ( Fig. 219 ); lateral lobes elongated and folded; stretcher and pit small ( Fig. 220 ). Median part of scape wide and short; genital pores positioned slightly pass the base of the lateral lobes ( Figs 221, 222 ). Internal genitalia with an oval ventral receptacula with a large rectangular base ( Fig. 221 ). Other material examined: USA : California : 16km E Hammond, 1066m , 20.iii.1941 , 1Ƥ, S., D. Mulaik ( AMNH ); Ave 312, 0.8km E road 196, 29 .iii–04.iv.1979, pitfall, oak duff, 1Ƥ, D. Caroll ( CAS ); Kaweah Lake, 30.iv.1978 , manzanita leaf litter, 13, 04 .xii.1978, 13, moss on rocks, D. Caroll ( CAS ); Robies del Rio, Hastings National History Reserve, 13, Linsdale ( AMNH ); San Francisco, 1Ƥ ( CAS ); Stanford University, 1920-1921 , 13, J. Chamberlin ( AMNH ). Oregon : Imbler, 26.ix.1964 , 13, J., W. Ivie ( AMNH ). Utah : Dry Canyon, 22.ii.1932 , 1Ƥ, 02.iv.1941 , 1Ƥ, W. Ivie ( AMNH ); Emmigration Canyon, 24.ii.1940 , 2Ƥ, S., D. Mulaik ( AMNH ); Salt Lake City, Mill Creek Canyon, 2438m , 05.xi.1939 , 1Ƥ, D. Mulaik ( AMNH ); Storm Mountain, Bog Cottonwood Canyon, 22.x.1939 , 13, W. Ivie ( AMNH ). Washington : Cow Lake [ 47.135N , 118.158W ] 545m , 01.vi.1991 , under rock at base of rim rock, 1Ƥ, R. Crawford ( UWBM ); Milk Canyon [ 46.855N , 120.819W ] 883– 914m ., 0 9–11.xi.1996 , under stones, 73, Y . Marusik ( AMNH ); Naches River [ 46.631N , 120.59W ] 359m , 23.ii.1991 , alder litter by road, 1Ƥ, R. Crawford ( UWBM ); Oak Creek [ 46.723N , 120.815W ] 524–549m , 29.ii.1992 , under rocks, 1Ƥ, R. Crawford ( UWBM ); Pitcher Canyon [ 47.365N , 120.348W ] 19.iv.1986 , under rocks, 1Ƥ, R. Crawford ( UWBM ); Wenatchee [ 47.438N , 120.357W ] 10.x.1980 , under rock, semi-dry, 23, 14 .x.1980, tall grass and stones, 1Ƥ, 20.x.1980 , maple duff, 23, D. Caroll ( CAS ); Wenatchee canyon [ 47.406N , 120.79W ] 27.x.1984 , damp gully, sparse litter, 132Ƥ, D. Caroll ( CAS ); Wenatchee, TFRC [ 47.438N , 120.357W ] 22.x.1982 , maple duff, 132Ƥ, 18.x.1984 , grass and maple duff, 832Ƥ, 31.iii.1985 , ravine, damp grass and duff, 137Ƥ, D. Caroll ( CAS ). Distribution: Western USA . FIGURES 213–222. Agyneta vinki n. sp. 213–217, Male. 218–222, Female. 213. Palp, retrolateral view. 214. Palp, prolateral view. 215. Embolus, prolateral view (arrow points to serrated extension). 216. Radical division, retrolateral view. 217. 218. Abdomen, dorsal view. 219. Epigynum, ventral view. 220. Internal genitalia, ventral view. 221. Internal genitalia, dorsal view. 222. Internal genitalia, lateral view (arrow points to basally folded lateral lobe). Agyneta panthera new species Figs 223–232 , map 15 Type material: Male holotype form California, Los Angeles Basin, Westwood, 12 Nov. 1952 , R. Schick ( AMNH ). EXAMINED . Etymology: The specific epithet is in reference to the coloration pattern of the legs ressembling a panthera . Diagnosis: Males can be separated from all Agyneta species by their greatly enlarged and strongly pointed ventral cymbial tubercle ( Fig. 224 ). Females are distinguished from most species by their wide proximal part of scape slightly expanding basally ( Fig. 229 ). To differentiate from A. vinki , see the diagnosis of the latter. Description: Male : Total length 1.79; carapace length 0.85, width 0.73. CEPHALOTHORAX: Carapace dark orange-brown, shiny, finely reticulate; suffused with dark gray along radiating lines and margin; trident mark present. Sternum dark brown, suffused with dark gray. Clypeus height 2. Chelicerae orange, with transverse dark gray suffused band, excavated; ~13 seta-tipped tubercles; promargin four teeth, retromargin three denticles, both margins with small rounded projection near base of fang. Cheliceral stridulatory organ ~32 striae, well spaced apically, narrowing basally. ABDOMEN: Patterned, with vertical gray band reaching halfway on the abdomen and five transverse gray chevrons ( Fig. 227 ). LEGS: Light orange, base and tip of femur, tibia and metatarsus with dark gray bands; leg I total length: 3.59; leg III total length: 2.28; Tm I: 0.26, Tm IV: absent. GENITALIA: Palpal retrolateral tibial apophysis small, pointed; dorsal tibial apophysis reduced, rugose; two retrolateral trichobothria and one dorsal ( Fig. 223 ). Cymbium triangular; glabrous depression present ( Fig. 223 ); dorsal cymbial tubercle absent; ventral cymbial tubercle elongated, smooth; prolateral notch shallow ( Fig. 224 ). Paracymbium apical pocket long, anterior pocket short and curved, making a medium cover, posterior pocket long and curved ( Fig. 223 ). Embolus tip pointed, with pointed projection; basally with one large prong; ventrally with numerous small spines; Fickert’s gland absent; ventral lamella transparent, small; thumb going slightly over the embolus proper ( Fig. 225 ). Embolus proper set apically, ventral part larger, not serrated ( Fig. 225 ). Anterior terminal apophysis with pointed tip and numerous long protrusions; posterior terminal apophysis with small squared tip; lamella characteristica large, with two pointed prongs and a blunt one ( Fig. 226 ). Female : Total length 2.52; carapace length 0.94, width 0.79. MAP. 15. Localities of Agyneta panthera n. sp. FIGURES 223–232. Agyneta panthera n. sp. 223–227, Male. 228–232, Female. 223. Palp, retrolateral view. 224. Palp, prolateral view. 225. Embolus, prolateral view. 226. Radical division, retrolateral view. 227. 228. Abdomen, dorsal view. 229. Epigynum, ventral view. 230. Internal genitalia, ventral view. 231. Internal genitalia, dorsal view. 232. Internal genitalia, lateral view. CEPHALOTHORAX: Same coloration as male. Chelicerae orange; promargin five teeth, retromargin four denticles. Cheliceral stridulatory organ ~50 striae, well spaced apically, narrowing basally. ABDOMEN: Patterned, same as male ( Fig. 228 ). LEGS: Same as male; palp tarsus normal, claw absent; leg I total length: 3.78; leg III total length: 2.67; Tm I: 0.23, Tm IV: absent. GENITALIA: Epigynum with wide proximal part of scape, very slightly enlarging apically; epigynal slits oval; pit hook depression minuscule ( Fig. 229 ); lateral lobes long and folded; stretcher medium; pit deep ( Fig. 230 ). Median part of scape long and narrow; genital pores situated in middle of the lateral lobes ( Fig. 231 ). Internal genitalia with elongated receptacula with large base ( Fig. 230 ). Other material examined: USA : Arizona : Grand Canyon, Colorado river side, mile 169, 27 .x.1982, Tamarix duff, 13, V. Roth ( AMNH ); Grand Canyon, Colorado river side, mile 172, 27 .x.1982, Tamarix duff, 13, V. Roth ( CAS ); North Hollywood, 01.xii.1979 , avocado litter, 23, D. Caroll ( CAS ). California : 24km W Santa Monica, 20.iii.1941 , 13, W. Ivie ( AMNH ); 9km SW Victorville, 09.iii.1941 , 1Ƥ, A., W. Ivie ( AMNH ); Hermosa Beach, 18.iii.1941 , 131Ƥ, W. Ivie ( AMNH ); Rancho Sante Fe, 01–14.1959, Alfafa field, 13, E. Schlinger ( AMNH ); San Joachim Valley, 16–31. vii.1995, 1 Ƥ, R. Schick ( AMNH ); Sunland, v.1936, 13 , R. Schick ( AMNH ); Topanga Canyon, 18.iii.1941 , 538Ƥ, W. Ivie ( AMNH ); Westwood, Los Angeles Basin, 12.xi.1952 , 13, R. Schick ( AMNH ). Nevada : Mercury, N.T.S, 07.iii.1961 , 13, 29.xi.1961 , 13 ( AMNH ). Distribution: Southwestern USA .