Revision and cladistic analysis of the Southeast Asian leaf-dwelling spider genus Calapnita Simon (Araneae, Pholcidae) Author Bernhard A. Huber text Zootaxa 2017 4219 1 1 63 journal article 37319 10.5281/zenodo.273086 0d2332a5-1968-49d7-a570-f31d386eccb3 1175-5326 273086 0FA0F51A-3868-4F13-A93D-E34CA5A689F8 Calapnita bariengi sp. nov. Figs 142–155 Diagnosis . Distinguished from most other species of vermiformis group (except C. bario ) by distal cheliceral apophyses not bipartite ( Figs 146, 154 ) and by shape of procursus (sclerotized ventral flap pointed dorso-distally, very simple dorsal part; Figs 142–143, 150–151 ); from several species also by narrow and distinct epigynal ‘knob’ ( Fig. 148 ), and by oval pore plates converging anteriorly and close together ( Fig. 149 ). Etymology . The species is named for Tun Datuk Patinggi Tan Sri Temenggong Jugah anak Barieng ( 1903– 1981 ), long-time chief of the Iban people, widely known for his statement “Do not be like the sugar cane, sweet at the beginning but tasteless at the end”. Material examined. Holotype . MALAYSIA-BORNEO: ♂ , ZFMK ( Ar 16023), Sarawak , Niah Cave National Park , forest near headquarters ( 3.820°N , 113.763°E ), 40 m a.s.l. , night collecting, undersides of leaves, 28.vii.2014 ( B.A. Huber , S.B. Huber ). Other material . MALAYSIA-BORNEO: 1♂ 3♀, ZFMK (Ar 16024), same data as holotype. Assigned tentatively . MALAYSIA-BORNEO: 1♂ , ZFMK ( Ar 16025), Sarawak , near Kapit , forest along river (1.937– 1.944°N , 112.904– 112.910°E ), 80–120 m a.s.l. , underside of leaf, 20.vii.2014 ( B.A. Huber , S.B. Huber ) ; 1♂ in absolute ethanol, ZFMK ( Bor 186), same data . Description. Male ( holotype ) MEASUREMENTS. Total body length 4.6, carapace width 0.65. Leg 1: 29.9 (7.7 + 0.3 + 6.6 + 12.8 + 2.5), tibia 2: 5.1, tibia 3: 3.1, tibia 4: 5.6; tibia 1 L/d: 120. Distance PME-PME 180 µm, diameter PME 90 µm, distance PME- ALE ~30 µm; no trace of AME. COLOR. Entire animal pale gray to whitish, legs slightly yellowish with brown patellae and tibia-metatarsus joints. BODY. Habitus as in close relatives (see e.g., Figs 120, 122 ); ocular area barely elevated, each triad on very low hump; carapace without median furrow; clypeus unmodified; sternum slightly wider than long (0.46/0.44), unmodified. CHELICERAE. As in Figs 146–147 , apophyses near lamellae not bipartite, pair of slender lateral processes; without modified hairs; without stridulatory ridges. PALPS. In general similar to C. vermiformis (cf. figs 139 and 140 in Huber 2011 ); trochanter apophysis as in Fig. 145 , with tiny tubercles and distinct subdistal process; femur as in Fig. 144 , distal process at 49% of femur length; tibia length/diameter 0.55/0.26; procursus as in Figs 142–143 , with simple ventro-distal flat sclerite; bulb length 0.32; embolus length 0.60. FIGURES 142–155 . Calapnita bariengi sp. nov. from Niah (142–149; ZFMK, Ar 16024) and from Kapit (150–155; ZFMK, Ar 16025). 142–143, 150–151 . Left procursi, prolateral and retrolateral views. 144, 152 . Left palpal femora, prolateral views. 145, 153 . Left palpal trochanters, retrolateral views. 146–147, 154–155 . Right male chelicerae, frontal and lateral views. 148 . Epigynum, ventral view. 149 . Cleared female genitalia, dorsal view. Scale lines: 0.5 mm (142–143, 150–151), 0.3 mm (144– 149, 152–155). LEGS. Without spines and curved hairs; few vertical hairs; retrolateral trichobothrium on tibia 1 at 2%; prolateral trichobothrium absent on tibia 1, present on other tibiae; tarsus 1 pseudosegments not visible in dissecting microscope. Male (variation). Tibia 1 in other male from type locality: 6.4. The specimens from Kapit are slightly larger ( 1♂ measured: leg tibia 1: 7.0; palpal tibia length/diameter 0.61/0.30; bulb length 0.42; embolus length 0.73), and differ slightly in shapes of male palps and chelicerae ( Figs 150–155 ); they are therefore assigned tentatively. Female . In general similar to male; eye triads slightly closer together (distance PME-PME 155 µm). Tibia 1 in 4 females : 5.4, 5.6, 6.0, 6.2. Epigynum as in Fig. 148 , weakly sclerotized triangular plate with small and slender anterior ‘knob’; internal genitalia as in Fig. 149 , with pore plates close together. ALS with only two spigots each (checked with light microscope). Distribution . Known from two localities in Sarawak ( Fig. 283 ).