Morphological and molecular characterization of Klossnema viguerasi n. sp (Nematoda: Oxyuridomorpha: Hystrignathidae) from a Cuban passalid beetle (Coleoptera: Passalidae), first record of the genus for Cuba Author Morffe, Jans 0000-0002-9968-8129 Instituto de Ecología y Sistemática, Carretera Varona 11835 e / Oriente y Lindero, La Habana 19, CP 11900, Calabazar, Boyeros, La Habana, Cuba. & Department of Environmental Biology, College of Bioscience & Biotechnology, Chubu University, 1200 Matsumoto, Kasugai, Aichi 487 - 8501, Japan. koichihasegawa @ isc. chubu. ac. jp; https: // orcid. org / 0000 - 0002 - 9968 - 8129 koichihasegawa@isc.chubu.ac.jp Author García, Nayla 0000-0002-3979-8086 Instituto de Ecología y Sistemática, Carretera Varona 11835 e / Oriente y Lindero, La Habana 19, CP 11900, Calabazar, Boyeros, La Habana, Cuba. & nayla @ ecologia. cu; https: // orcid. org / 0000 - 0002 - 3979 - 8086 nayla@ecologia.cu Author Hasegawa, Koichi 0000-0002-9968-8129 Department of Environmental Biology, College of Bioscience & Biotechnology, Chubu University, 1200 Matsumoto, Kasugai, Aichi 487 - 8501, Japan. koichihasegawa @ isc. chubu. ac. jp; https: // orcid. org / 0000 - 0002 - 9968 - 8129 koichihasegawa@isc.chubu.ac.jp Author Breugelmans, Karin 0000-0002-1236-7403 Royal Belgian Institute of Natural Sciences, Vautierstraat 29, B- 1000, Brussels, Belgium. karin. breugelmans @ naturalsciences. be; https: // orcid. org / 0000 - 0002 - 1236 - 7403 karin.breugelmans@naturalsciences.be text Zootaxa 2021 2021-02-22 4933 3 391 402 journal article 7977 10.11646/zootaxa.4933.3.6 9825c147-58b8-40c9-bc7f-ad34a7b90991 1175-5326 4555057 BFA44668-69F6-435B-871A-5DB85D490710 Klossnema Cordeiro & Artigas, 1983 Emended diagnosis. General. Small and slender nematodes. Cephalic capsule smooth, dorsoventrally compressed. Mouth hexagonal, laterally orientated, surrounded by six labia, each of them coinciding with one of its sides. Labia set-off from each other by short cleavages coinciding with mouth edges. Four flat, elongated cephalic papillae present, arranged as one dorsal pair and one ventral. Papillae of each pair touch dorsally and ventrally, respectively, forming obtuse angles. Digitiform structure present close to each lateral edge of the mouth. Cuticle unarmed and finely annulated from the base of the cephalic capsule to the posterior region. Lateral alae absent. Oesophagus with a muscular, sub-cylindrical procorpus, its base slightly dilated. Isthmus comparatively long. Basal bulb pyriform, valve plate well developed. Intestine simple, sub-rectilinear, anterior portion barely dilated. Nerve ring encircling procorpus at its posterior half. Excretory pore ventral, post-bulbar. Female. Reproductive system monodelphic-prodelphic. Eggs comparatively large, ovoid, smooth-shelled. Tail very short, subulate and curved, with a hook-like appearance. Male. Body smaller and slightly less robust than females, posterior end ventrally curved. Monorchic. Spicule absent. Tail very short, conoid, its tip sharp. Dorsal cuticle of the tail end thickened and smooth. Six copulatory papillae present, four pre-cloacal and two post-cloacal. First pre-cloacal pair consists of a ventromedian duplex papilla on a protuberance, the sensilla of each papilla of this pair surrounded by peg-like prominences, arranged in more or less concentric circles. Second pair of pre-cloacal papillae formed by large peg-like papillae, lateral in position, located at a short distance before the level of the cloaca. The pair of post-cloacal papillae consists of sub-lateral minute papillae near the tail tip. Phasmids pore-like, located at the tail tip.