Cladistic analysis and taxonomic revision of the genus Karos Goodnight & Goodnight, 1944 (Opiliones, Laniatores, Stygnopsidae) Author Cruz-López, Jesús A. Author Francke, Oscar F. text Zoological Journal of the Linnean Society 2015 2015-11-18 175 4 827 891 http://dx.doi.org/10.1111/zoj.12299 journal article 10.1111/zoj.12299 0024-4082 5339971 CHAPULOBUNUS POBLANO SP. NOV. ( FIGS 32–34 , 58 , 60H , 61G , 63A, J, K , 64K ) Type material: MEXICO : Puebla : ♂ holotype , and 1 ♂ and 3 ♀ paratypes [ CNAN-T0795 and CNAN- T0796 ( 1.ix.2009 ; O. Francke and A. Valdez )] ( 19°51′56.80″N , 97°33′05.58″W ). Municipio Zacapoaxtla , Estación de microondas Tomaquillo . Material examined: MEXICO : Puebla : 1 ♀ [ CNAN ( 30.ix.2006 ; O. Francke , A. Valdez , and H. Montaño )] ( 19°48′46.80″N , 97°25′0.37″W ). Municipio Atempan , 1 km SE of Tacopan. Hidalgo : 6 ♂ and 6 ♀ [ CNAN ( 23.ix.2006 ; O. Francke , A. Valdez , and H. Montaño )], ( 20°8′30.01″N , 98°10′44.4″W ). Municipio Acaxochitlán , 2 km E of Acaxochitlán. Etymology: The specific epithet is a noun in apposition, derived from the Spanish word poblano , which means originating from the state of Puebla , Mexico . Diagnosis: Dark brown species. Ocularium spiniform, with long median spine ( Fig. 32 ). Ventral ornamentation of leg IV in males consists of big spiniform tubercles, two mesal on femur and five to six along tibia ( Fig. 33A ). Males with bulge on anal plate ( Fig. 63A ). Male genitalia: with six to nine pairs of lateral setae, three laterobasal pairs of parastylar setae. Lateral projection of follis thin, pointed distally ( Fig. 34 ). Figure 32. Chapulobunus poblano sp. nov. A, habitus, dorsal view. B, scutum, dorsal view. C, ocularium, frontal view. Scale bars: A = 5 mm, B = 2.5 mm, C = 1 mm. Figure 33. Chapulobunus poblano sp. nov. A, legs IV, mesal view. B, habitus, ventral view. C, habitus, lateral view. Scale bars: A = 3.3 mm, B = 2.5 mm, C = 2 mm. The dark lines on (B) indicate the stigmatic area. Description Male holotype : Measurements : scutum length 4.4, maximum scutum width 4.2. Legs. I 1.90/0.65/1.40/ 1.65, II 2.90 /1.00/2.20/2.35, III 2.70 /1.00/2.00/2.25, IV 3.45 /1.25/3.25/3.45. Dorsum . Scutum covered by nu- merous rounded tubercles ( Fig. 32 ). Ocularium with a acute median spine ( Fig. 32C ). Extra row of pegs composed of six to seven pegs, sometimes bifid ( Fig. 32B ). Spine on area II lower than the ocularium, spine on area III with similar height to ocularium, noticeably bent backwards ( Fig. 33C ). Sulcus between areas III and IV almost absent, vestigial. Lateral clear areas on scutum projected in semi-teardrop shape, project- ed in apices of area V and free tergites I and II , teardrop-shaped, absent in free tergite III . Free tergites with median row of tubercles. Venter. Finely covered by small, rounded tubercles. Stigmatic area inverse ‘Y’ shaped ( Fig. 33B ). Coxa IV with one dorsoectal spiniform tooth. Ornamentation of the anal plate heterogeneous , 862 J. A. CRUZ-LÓPEZ AND O. F. FRANCKE Figure 34. Chapulobunus poblano sp. nov. , male genitalia. A, dorsal view. B, ventral view. C, lateral view. White arrows indicate ventral microsetae in (B) and parastylar setae in (C). with median bulge rounded ( Fig. 63A ). Pedipalps. Patella with two mesal spiniform tubercles. Legs. I and II noticeably thinner and less ornate than the posterior legs. Femora III and IV curved, with ventral ornamentation increasing in size distally. Femur with two spiniform tubercles on ventromesal side, tibia with six or seven spiniform tubercles along the ventral side. Femur and tibia IV swollen ( Fig. 33A ). Tarsal count 4(2):5(2):6:6. Male genitalia. Pars distalis swollen in the middle, apical margin convex, rounded. Six to nine pairs of lateral setae. Three pairs of parastylar setae laterobasal to follis. Spiniform projections noticeably exposed. Four to five pairs of ventral microsetae, forming an irregular row along ventrolateral margin of pars distalis. Lateral projection of follis lobular, pointed distally ( Fig. 34 ). Female: Measurements: scutum length 4.3, maximum scutum width 4.1. Similar to male, with the following differences: ventral armature and thickness of femur and tibia IV reduced ( Fig. 33A ). Tarsal count 4(2):5/6(3):5/6:6. Natural history: Similar to Chapulobunus unispinosus .