Morphology-based cladistics splinters the century-old dichotomy of the pied harvestmen (Arachnida: Gonyleptoidea: Cosmetidae) Author Medrano, Miguel Author Kury, Adriano Brilhante Author Mendes, Amanda Cruz text Zoological Journal of the Linnean Society 2022 2021-08-30 195 585 672 journal article 113070 10.1093/zoolinnean/zlab043 e070d52f-d008-4f08-988a-a8b17c27418f 0024-4082 6959516 1C89556A-12CB-43B7-9B49-E02EFF1543D3 PROTUS BOLIVARI ( ROEWER, 1952 ) COMB. NOV. ( FIGS 1G, H , 30–32 ) Cynortula bolivari Roewer, 1952: 43 , fig. 26. Type data: Holotype , Peru . San Martín : Juanjui , Huallaga River above 350 m , [–7.20416°, –76.73819°, 350 m ], 1 ♂ ( SMF RII 9813 /277, not examined) . Paratypes : Same data as holotype 1 ♂ ( WPC , not examined) Non-type material: Peru . [ San Martín ]: Juanjui , Huallaga River , [–7.20416°, –76.73819°, 350 m ], Ortiz & Beltran leg., examined by photograph, 1 ♀ ( AMNH ). Madre de Dios : distrito de Tambopata , Reserva Nacional de Tambopata [–12.82822°, –69.29056°], 1 ♂ (USNMENT 511638); [–12.52339°, –71.18131°], 1 ♂ (USNMENT 511626); Zona reservada Tambopata [–12.83333°, –69.28333°, 290 m ], J Coddington leg., 7.vi.1988 , (post-fire material) 1 ♂ (USNMENT 1538074); 8.vi.1988 , (post-fire material) 1 ♀ (USNMENT 1538075) . Diagnosis: It differs from the other members of Protus by having area III armed with larger paramedian yellow tubercles than those of area II ( Fig. 31A–C ), DS with scattered yellow dots in abdomen and carapace ( Fig. 31A ) and by the combination of the following features: carapace with a thin, yellow stripe bordering anterior margin of carapace ( Fig. 31A ), VP of the penis not elongated and with distal border substraight ( Fig. 32C ), stylus with short wattle ( Fig. 32D ).