Review of Oculogaster Kluge 2016 (Ephemeroptera, Baetidae, Procloeon Bengtsson 1915) Author Kluge, Nikita J. text Zootaxa 2020 2020-07-29 4820 3 401 437 journal article 8958 10.11646/zootaxa.4820.3.1 fd413580-2960-463f-a924-8c27eaeafc6e 1175-5326 4397861 44BD1E07-C9F3-4488-936E-819C2FCA18C8 Oculogaster Kluge 2016 ( Figs 1–133 ) Subgenus Oculogaster Kluge 2016: 494 . Type species: Procloeon cylindroculum Kimmins 1956 . Diagnosis. (1) Viviparous ( Kluge 2016 : figs 38–42, 76–79). The same in Cloeon s. str. and some other mayflies, but not in other Procloeon /g1. (2) Pterostigma normally either with 1 cross vein ( Fig. 40 ), or with 2 cross veins ( Fig. 104 ) (on individual wings this number varies from 1 to 3) ( Figs 40–41 ). (3) Larval femur with two apical spine-like setae ( Figs 17 , 81 , 110 ). The same in many other taxa of Baetidae , but not in other Procloeon /g1 and not in other Cloeon /fg1. Larvae of all Oculogaster have a similar structure of the labium, with a pair of submedian setae distinguishable among other setae of dorsal surface ( Figs 50 , 113 ). Mandibles of the « Baetis - type » ( Kluge 2020a : table 1 and p. 578), i.e. with incisor and kinetodontium fused, left prostheca wide, and all denticles of incisor, kinetodontium, prostheca and mola visible in one view ( Figs 51–52 , 114–115 ). All species of Oculogaster lack hind wings, and larvae have no vestiges of hind protoptera ( Müller-Liebenau & Hubbard 1985 : fig. 9h). Distribution. Paleotropical (Afrotropical and Oriental Regions). Discussion. In the previous diagnosis, the number of cross veins in pterostigma was reported as one, with individual variability; this diagnosis was based on P. ( O. ) cylindroculum and P. ( O. ) album , whose pterostigma normally has one cross vein. Examination of P. regularum reveals that it should be placed in Oculogaster , while its pterostigma normally has 2 cross veins; this fact expands the diagnosis of Oculogaster . While pterostigma with one cross vein is not found in any taxa other than Oculogaster , pterostigma with two veins occurs in some other taxa, particularly in some species of Cloeon s. str. , which were confused with species belonging to Oculogaster . Species composition and classification. Oculogaster can be divided into two species groups: the African group, which includes P. ( O. ) cylindroculum Kimmins 1956 , P. ( O. ) barnardi sp. n. , P. ( O. ) niger sp. n. , P. ( O. ) sp. «Wolfkloof» and presumably P. ( O. ) silvicola Gillies 1997 , and the Asian group, which includes P. ( O. ) album Kluge 2016 , P. ( O. ) regularum Müller-Liebenau & Hubbard 1985 , P. ( O. ) malabarensis sp. n. , presumably P. julia ( Gillies 1949 ) comb. n. and unnamed P. sp. «Thailand-3», P. sp. «Sulawesi» and P . sp. « Philippines ».