Taxonomy and evolution of asymmetric male genitalia in the subgenus Ashima Chen (Diptera: Drosophilidae: Phortica Schiner), with descriptions of seven new species Author Toda, Masanori J. Hokkaido University Museum, Hokkaido University, Sapporo 060 - 0810, Japan Author Bänziger, Hans Department of Entomology and Plant Pathology, Faculty of Agriculture, Chiang Mai University, Chiang Mai 50200, Thailand hans. banziger @ cmu. ac. th; https: // orcid. org / 0000 - 0001 - 8913 - 3034 Author Sati, Pradeep C. Systematic & Cytogenetic Laboratory, Department of Zoology and Biotechnology, HNB Garhwal University, Srinagar-Garhwal, 246174, Uttarakhand, India. & satipradeep 28 @ gmail. com; https: // orcid. org / 0000 - 0001 - 6809 - 8575 Author Fartyal, Rajendra S. Systematic & Cytogenetic Laboratory, Department of Zoology and Biotechnology, HNB Garhwal University, Srinagar-Garhwal, 246174, Uttarakhand, India. & rs. fartyal @ hnbgu. ac. in; https: // orcid. org / 0000 - 0002 - 8844 - 6263 Author Suwito, Awit Zoology Division (Museum Zoologicum Bogoriense), Research Center for Biology-LIPI, Jl. Raya Jakarta-Bogor Km. 46, Cibinong 16911, West Java, Indonesia. awitswt @ yahoo. com Author Katoh, Toru Department of Biological Sciences, Faculty of Science, Hokkaido University, Sapporo 060 - 0810, Japan t _ katoh @ sci. hokudai. ac. jp; https: // orcid. org / 0000 - 0003 - 3376 - 8545 text Zootaxa 2020 2020-06-08 4789 1 1 54 journal article 21673 10.11646/zootaxa.4789.1.1 c12aaed8-264c-459c-b1aa-09eaeb686670 1175-5326 3884610 311A27A2-A053-4D20-A43E-EF57DBE55166 2.2. Phortica ( Ashima ) tanabai species subgroup, new Diagnosis. Prementum with a pair of patches of pubescence (ch.12-1; Fig. 6B ); aedeagal, outer membrane ornamented with somewhat long, conical spines (ch.55-0, ch.56-2; Figs 6E,G , 7G,H ). Shared characters. Some interfrontal setulae as long as lowest setulae on ocellar triangle (ch.3-0; Fig. 7B ). Arista apically with leaf-like expansion (ch.7-1; Figs 6A , 7C ). Setulae just before prescutellar acrostichal setae somewhat longer than other acrostichal setulae (ch.15-1). Fore and mid tibiae entirely yellow (ch.17,18-0). Abdominal sternites III–V wider than long (ch.22,24,25-1; Figs 6C , 7D ); III widest among sternites (ch.23-1; Figs 6C , 7D ), with dense setulae on medial surface (ch.26-1; Figs 6C , 7D ). Pregonite not concave on dorsosubapical margin of apical knob (ch.48-0; Figs 6E , 7G ), without basal extension (ch.49-0; Figs 6E , 7G ). Posterior, rod-like process of phallapodeme not longer than aedeagal median rod (ch.62-0; Fig. 6E ). Aedeagal sheath without lateral lobes bearing strongly sclerotized postgonites (ch.65-0; Figs 6F , 7F ). Included species. Phortica ( Ashima ) huiluoi Cheng & Chen *; P . ( A .) mediarista ; P . ( A .) tanabei ; P . ( A .) akutsui sp. nov. ; and P . ( A .) halimunensis sp. nov. * The assignment was judged from the original description by Cheng et al . (2008) .