Macrodactylini (Coleoptera, Scarabaeidae, Melolonthinae): primary types of type species and taxonomic changes to the generic classification Author Fuhrmann, Juares Author Vaz-De-Mello, Fernando Z. text European Journal of Taxonomy 2017 2017-09-13 350 1 71 journal article 22059 10.5852/ejt.2017.350 52e4ae01-f51f-4c4a-8a14-9fb5b17788d6 2118-9773 3832682 8D14DBDE-AD13-445B-B2D0-221F19FC7C37 Chariodactylus Moser, 1919 Type species Chariodactylus chacoensis Moser, 1919 (designation: Evans 2003: 247 ). Diagnosis Clypeus and frons gradually raised over the lateral area of head ( Fig. 11A ); clypeal anteroventral area strongly reduced (similar to Fig. 11H ); posterior ocular area exposed ( Fig. 11A ), eye large; pronotum as long as wide, with anterior and posterior margins not beaded; prosternum with two anterior sulci (similar to Fig. 11H ); protibia with two external teeth and without spur ( Fig. 11A ); abdomen with intersegmental membrane VII–VIII exposed. Remarks The genera Chariodactylus , Macrodactylus and Manodactylus Moser, 1919 are similar to each other. Chariodactylus ( Fig. 11A ) is distinguished as follows (opposition to 1, Macrodactylus ( Fig. 11C ); 2, Manodactylus ( Fig. 11B )): clypeus broad, trapezoid (1, narrow, trapezoid; 2, as Chariodactylus ); eye large (1, 2, small); pronotum wider than long (1, longer than large; 2, as Chariodactylus ). Future cladistic analysis and a taxonomic review of Macrodactylus including Chariodactylus and Manodactylus are needed to confirm the possible synonymy between these genera.