Revision of Holarctic Teleiodini (Lepidoptera: Gelechiidae) Author Lee, Sangmi Author Brown, Richard L. text Zootaxa 2008 1818 1 55 journal article 10.5281/zenodo.182949 834831a3-c0b8-44fa-b357-c14c05822909 1175-5326 182949 Agnippe Chambers Agnippe Chambers, 1872b : 194 . Type species: Agnippe biscolorella Chambers, 1872 , by subsequent designation ( Meyrick 1925 ). Aganippe Chambers, 1880b : 198 . Misspelling of Agnippe Chambers, 1872 . Phaetusa Chambers, 1875f : 105 . Preoccupied by Phaetusa Wagler, 1832 . Evippe Chambers, 1873a : 185 . New Synonymy Type species: Evippe prunifoliella Chambers, 1873 , by monotypy. Tholerostola Meyrick, 1917 : 40. Type species: Tholerostola omphalopa Meyrick, 1917 , by monotypy. Description . Imago ( Figs. 51–52 ). Labial palpus long, extending beyond vertex, second and third segments subequal in length, second segment slightly thickened with scales dorsoventrally. Antenna longer than half forewing length. Clypeus with ventral margin rounded. Ocellus absent. Posterior area of sitophore with four campaniform sensilla in line except one or other of middle two offset; anterior area with four campaniform sensilla. Forewing (length/width ratio 4.1) without tufts of raised scales, R1 absent in male, present in female, R5 and M1 stalked, M2 and M3 stalked, CuA1 arising halfway between M3 and CuA2 ( Fig. 10 ); median fascia present or absent, if present, directed from base of costa toward posterior margin, often with sinuate line separating dark costal area and pale posterior area. Hindwing narrower than forewing (length/width ratio 4.6), M1 absent, M2 close to M3, M3 and CuA1 separate ( Fig. 15 ), male with hair pencil present at base of anal area or at 1/3 costa. Male abdominal tergum VIII lingulate, with anterolateral hair pencil; sternum VIII broad, anterior margin broadly emarginate. Female abdominal segment VIII evenly sclerotized, without special modifications. Male genitalia ( Fig. 25 ): uncus elongate, slender, rounded distally, with slight mesial incision, subequal in length with gnathos; gnathos with spatulate dorsal part and trifid apex of ventral part; costal part of valva as long as length of tegumen, digitate with small apical dentate projection; saccular part of valva lobelike; tegumen basal width/length ratio 0.8; vinculum with pair of weak mesial projections; saccus as long as length of gnathos, rodlike; phallus stout, some species with group of needlelike cornuti. Female genitalia ( Fig. 40 ): apophyses posteriores 2 x length of apophyses anteriores; apophyses anteriores subequal in length with abdominal segment VIII; antrum broadly funnel shaped to tubular; ductus bursae broad, shorter than species in Teleiodes ; distinct accessory bursae with row of well-developed spinules; signum suboval with sinuous posterior margin, irregular anterior edge and with transverse ridge. Larva. Undescribed. Pupa. Body smooth; abdominal segment VII with lateral cavity bordered by fringe of setae and with cephalic margin trilobed ( Mosher 1916 ). Diagnosis. Agnippe and some species of Aristotelia have a similar wing pattern, but Agnippe differs by the forewing having R5 and M1 stalked and M2 and M3 stalked, and the male genitalia having a split valva and a dorsal part of gnathos ( Fig. 25 ). Hosts. Fagaceae : Quercus spp. ( A. laudatella ). Fabaceae : Lespedeza sp. ( A. sp .), Gleditsia sp. ( A. biscolorella ). Rosaceae : Cercocarpus ledifolius Nutt. ( A. abdita ), Prunus spp. ( A. prunifoliella ). ( Forbes 1923 ; Kimball 1965 ; Okada 1961 ; Opler 1974 ; Robinson et al. 2002 ). Diversity and distribution. Agnippe includes 23 species occurring throughout North America , southeastern Europe, and Asia ( Amsel 1961 ; Forbes 1923 ; Huemer and Karsholt 1999 ; Keifer 1927 ; Li 1993 ; Li and Zheng 1998 ; Meyrick 1925 ; Okada 1961 ; Wang and Li 1994 ). Notes. Chambers (1873a) differentiated Evippe from Agnippe based on the absence of R 1 in the forewing (termed the first subcostal branch), but he mentioned his uncertainty in naming this genus. Although R1 is absent in the male of the type species, E. prunifoliella , this vein is present in the female as well as in the female of A. crinella Keifer. Busck (1906, fig. 1) illustrated a male forewing of A. evippeella Busck that lacks R1. Male and female genital characters indicate all species belong to one genus.