Genetic and morphological evidence for two species of Leucocarbo shag (Aves, Pelecaniformes, Phalacrocoracidae) from southern South Island of New Zealand Author Nicolas J. Rawlence Author R. Paul Scofield Author Hamish G. Spencer Author Chris Lalas Author Luke J. Easton Author Alan J. D. Tennyson Author Mark Adams Author Eric Pasquet Author Cody Fraser Author Jonathan M. Waters Author Martyn Kennedy text Zoological Journal of the Linnean Society 2016 177 676 694 journal article 10.1111/zoj.12376 b49e437b-c2a1-4e2a-a683-84c637a0535b 270312 LEUCOCARBO CHALCONOTUS (G. R. GRAY, 1845) OTAGO SHAG. Graucalus auritus Lesson, 1831 : G. R. Gray, 1843, in E. Dieffenbach, Travels in N.Z . 2: 201. Not Carbo auritus Lesson, 1831 . Gracalus chalconotus G. R. Gray, 1845 ; in Richardson & J. E. Gray (eds), Zool. Voy. Erebus & Terror: Birds 1(8): 20, pl. 21 – Otago Province, possibly Karitane (Scofield et al. , 2012). Graculus glaucus ‘Gray’: Reichenbach, 1850, Nov. Syn. Avium ; pl. 49, fig. 2553 – Otago. Phalacrocorax glaucus Hombron & Jacquinot, 1853 , in Dumont d’Urville, Voyage ˆ Sud, Zoologie 3: 127. Graculus chalconotus (G. R. Gray) : G. R. Gray, 1862, Ibis 4: 252. Graculus glaucus (Hombron & Jacquinot, 1853) : Bonaparte, 1855, Consp. Gen. Avium 2: 171. Phalacrocorax chalconotus (G. R. Gray) : Buller, 1873, Hist. Birds N.Z . (1st ed.), 334. Buller, 1888, Hist. Birds N.Z . (2nd ed.) 2: 162. Phalacrocorax carunculatus (Gmelin) : Buller, 1888, Hist. Birds N.Z . (2nd ed.) 2: 155. In part. Phalacrocorax chalconotus (G. R. Gray) : Buller, 1888, Hist. Birds N.Z . (2nd ed.) 2: 162. Phalacrocorax glaucus Hombron & Jacquinot : Buller, 1888, Hist. Birds N.Z . (2nd ed.) 2: 163. Phalacrocorax huttoni Buller, 1888 : Hist. Birds N.Z . (2nd ed.) 2: 174 – ocean beach near Dunedin, most likely Ocean Beach at Saint Clair, Dunedin (this study). Phalacrocorax chalconotus (G. R. Gray) : Ogilvie- Grant, 1898, Cat. Birds Brit. Mus. 26: 369. In part. Hypoleucus campbelli stewarti (Ogilvie-Grant) : Mathews & Iredale, 1913, Ibis 1 (10th Ser.): 413. In part. Hypoleucus chalconotus (G. R. Gray) : Mathews & Iredale, 1913, Ibis 1 (10th Ser.): 413. In part. Hypoleucus chalconotus (G. R. Gray) : Mathews, 1927, Syst. Avium Australasianarum 1: 227. In part. Phalacrocorax chalconotus (G. R. Gray) : Oliver, 1930, N.Z. Birds (1st ed.): 192. In part. Phalacrocorax huttoni Buller : Oliver, 1930, N.Z. Birds (1st ed.): 190. In part. Hypoleucus chalconotus (G. R. Gray) : Mathews, 1931, List Birds Australasia : 146. In part. Phalacrocorax ( Leucocarbo ) huttoni Buller : Falla, 1932, Rec. Auck. Inst. Mus. 1: 147. In part. Phalacrocorax ( Leucocarbo ) chalconotus (G. R. Gray) : Falla, 1932, Rec. Auck. Inst. Mus. 1: 148. In part. Hypoleucus chalconotus (G. R. Gray) : Mathews, 1936, Suppl. Birds Norfolk & Lord Howe Islands : 142. In part. Hypoleucus huttoni (Buller) : Mathews, 1936, Suppl. Birds Norfolk & Lord Howe Islands : 141. In part. Phalacrocorax ( Leucocarbo ) carunculatus chalconotus (G. R. Gray) : Checklist Committee Ornithological Society of New Zealand , 1953, Checklist N.Z. Birds : 29. In part. Leucocarbo carunculatus chalconotus (G. R. Gray) : Checklist Committee OSNZ, 1970, Annot. Checklist Birds N.Z. (2nd ed.): 32. In part. Leucocarbo chalconotus (G. R. Gray) : van Tets, 1976, Proc. 16th Int. Ornith. Cong .: 122. In part. Phalacrocorax carunculatus chalconotus (G. R. Gray) : Dorst & Mougin, 1979, in Peters, Check-list Birds World 1 (2nd ed.): 176. In part. Euleucocarbo chalconotus (G. R. Gray) : Siegel-Causey, 1988, Condor 90: 892. In part. Leucocarbo chalconotus (G. R. Gray) : Checklist Committee OSNZ, 1990, Checklist Birds N.Z. (3rd ed.): 83. In part. Phalacrocorax chalconotus (G. R. Gray) : O’Brien, 1990, in Marchant & Higgins, Handbook Aust. N.Z. Antarct. Birds : 876. In part. Leucocarbo ( carunculatus ) chalconotus (G. R. Gray) : Johnsgard, 1993, Cormorants, Darters and Pelicans World : 271. In part. Leucocarbo chalconotus (G. R. Gray) : Checklist Committee OSNZ, 2010, Checklist Birds N.Z. (4th ed.): 147. In part. Leucocarbo chalconotus ‘Otago’: Rawlence et al. , 2014, PLoS ONE 9: e90769: 1.