Two new species and two new combinations in Saphenista Walsingham, 1914 from western North America (Lepidoptera: Tortricidae) Author Brown, John W. text Insecta Mundi 2019 2019-12-03 738 738 1 8 journal article 10.5281/zenodo.3676593 f86beed9-5cc5-40cc-96c7-df36295b3e13 1942-1354 3676593 68DF38B0-C69E-4394-8992-CD74633ECADA Saphenista bartellae Brown , new species Fig. 1, 2 , 7, 11 Diagnosis. Saphenista bartellae is easily distinguished from all congeners by facies alone ( Fig. 1, 2 ), which are more similar to those of Aethes Billberg, 1820 than to other species of Saphenista . The forewing has a uniform white to cream ground color with a gray-brown, nearly complete median fascia of nearly uniform width, the costal portion of which is represented by a small, disjunct, semicircular patch near the mid-costa. Other North American congeners have a more mottled ground color and a variably developed, brownish median fascia from the hind margin that usually attenuates or terminates at or before the discal cell. In the male genitalia of S. bartellae ( Fig. 7 ) the spur from the vinculum is much longer and narrower than in other North American congeners. In the female genitalia of S . bartellae ( Fig. 11 ) the posterior margin of the sterigma is distinctly V-shaped with a strongly sclerotized edge, and this feature is shared with no other Saphenista . Description. Head . Vertex and frons white; labial palpus white, with scattered brown scales on second segment laterally and ventrally, scaling only slightly expanded distally on segment II, segment III exposed; antenna with white scales, sensory setae 2.0–2.5 times flagellomere diameter in male, extremely short, sparse in female. Thorax . Nota white to cream; scales of tegula rather short, white; no mid-posterior crest; forewing length 6.0–7.0 mm (mean = 6.5 mm, n = 10); forewing ground color white, sometimes with faint cream-ocherous overscaling, with a well-defined gray-brown median fascia extending from near mid-costal to middle of hind margin, interrupted subcostally, leaving a small gray-brown semicircular patch at costa; often with a rhomboidal gray-brown patch subapically, with a few extremely short, gray-brown strigulae scattered along costa from base to apex of forewing. Fringe concolorous with forewing. Hindwing white to pale cream with faint pale gray-brown overscaling; male with distinct costa roll (hair pencil) ca. 0.5 length of costa; frenulum with one spine in male, three in female (rarely two). Fringe whitish to pale cream. Abdomen . Lacking specialized processes. Mostly pale gray-cream. Male genitalia ( Fig. 7 ) with tegumen flattened at top; uncus absent; socii slender, erect, densely hairy, arising from lateral edge of top of tegumen; valva slender, gently upcurved throughout, widest at base, weakly attenuate throughout to narrow apex; sacculus short, confined to basal 0.1 of venter of valva, followed distally by a small lobe; transtilla well developed with median process bearing a wide, T-shaped expansion distally, lateral margins of process mostly parallel-sided throughout, only slightly expanded distally; vinculum arms joined distally, each bearing a long, slender flange, pointed distally; phallus large, almost as long as valva, slightly curved at ca. 0.6 distance from base, apically with slender ventral projection with blunt apex, vesica with a single slender cornutus. Female genitalia ( Fig. 11 ) with papillae anales slipper-shaped; apophyses unmodified, slender, length of apophyses anteriores about equal to that of apophyses posteriores; sterigma with posterior margin deeply V-shaped immediately posteriad of ostium, with strongly sclerotized margins; ductus bursae moderately wide, abruptly expanded ca. 0.3 distance from ostium to corpus bursae, with longitudinal wrinkles and membranous expansion on left side; corpus bursae short, ovoid, with sparsely distributed, minute spicules on entire inner surface, frail ductus of accessory bursae arising near middle of corpus. Types. Holotype ♂ , USA , Colorado , Gilpin Co. , 2538 Golden Gate Canyon Road (State Highway 46), 2674 m , 28 July 2016 , B. K. Bartell ( USNM ). Paratypes ( 41♂ , 28♀ ): CANADA : British Columbia : Vancouver, Stanley Park Aquarium, 15 m , 24 Jul 2007 ( 2♀ ), A. Li (CNC). USA : California : Kern Co.: Mt. Pinos, 6800′, 2 Jul 1965 ( 2♂ , 1♀ ), J. Powell (EME). Marin Co.: Inverness, Kehoe road, 75 m , 10–16 Oct 2001 ( 1♂ ), 17–23 Oct 2001 ( 1♂ ), 24–31 Oct 2001 ( 5♂ ), 1–6 Nov 2001 ( 2♂ ), J. Powell (EME). Inverness Park 15–22 Oct 1998 ( 1♀ ), 15–21 Oct 1999 ( 1♂ , 1♀ ), 20–26 Nov 2003 ( 2♂ ), J. Powell (EME). Inverness, top of Highland Road, 125 m , 9–16 Oct 2000 ( 1♂ ), 17–24 Oct 2000 ( 1♂ ), 25 Oct–2 Nov 2000 ( 1♂ , 2♀ ), 11–17 Nov 2004 ( 1♂ , 1♀ ), J. Powell (EME). W of Inverness, Kehoe Way, 75 m , 10–16 Oct 2001 ( 1♂ ), J. Powell (EME). Placer Co.: Ward Creek, 2 mi S Tahoe City, 6250’, 14 Jul 1984 ( 1♂ ), 16 Jul 1988 ( 1♀ ), 20 Jul 1988 ( 1♀ ), 27 Jul 1991 ( 2♂ ), 7 Aug 1983 ( 1♀ ), N. Westerland (LACM). Tuolomne Co.: Twain Harte, 21 Jul 1985 ( 1♂ ), M. Lundgren (EME). Colorado : Gilpin Co.: 2538 Golden Gate Canyon Road (State Highway 46), 2674 m , 12 Jul 2010 ( 1♂ ), 3 Aug 2010 ( 1♂ ), 12 Aug 2010 ( 1♀ ), 13 Jul 2012 ( 1♂ ), 17 Jul 2012 ( 1♂ ), 22 Jul 2014 ( 1♀ ), 23 Jul 2014 ( 1♂ ), 24 Jul 2014 ( 2♀ ), 17 Aug 2014 ( 1♂ ), 24 Aug 2015 ( 1♂ ), 26 Aug 2015 ( 1♂ ), 14 Jul 2016 ( 1♂ ), 16 Jul 2016 ( 1♀ ), 17 Jul 2016 ( 1♀ ), 18 Jul 2016 ( 1♂ ), 31 Jul 2016 ( 1♂ ), 2 Aug 2016 ( 1♀ ), 3 Aug 2016 ( 1♂ , 1♀ ), 8 Aug 2016 ( 1♂ , 1♀ ), 10 Aug 2016 ( 1♀ ), 11 Aug 2016 ( 1♂ ), B. K. Bartell (CSU). 2538 Golden Gate Canyon Road (State Highway 46), 2674 m , 11 Jul 2010 ( 1♀ ), 8 Aug 2010 ( 2♀ ), 10 Aug 2010 ( 1♂ ), 22 Jul 2011 ( 1♀ ), 27 Jul 2010 ( 1♂ ), 13 Aug 2011 ( 1♀ ), 14 Aug 2010 ( 1♂ ), 18 Aug 2011 ( 1♀ ), B. K. Bartell (DMNS). La Plata Co.: San Juan National Forest, 0.5 mi N Chris campground, 8 mi N Hermosa, 30 Aug 1991 ( 1♀ ), P. Opler (CSU). Montana : Mineral Co.: Superior, 4 Jul 1958 ( 1♂ ), M. May (USNM). Distribution and biology. Saphenista bartellae is recorded from coastal California (Marin Co.) (EME), north to Vancouver, British Columbia (CNC), at elevations of 15–75 m , east to the Rocky Mountains in Colorado and Montana at elevations of 2600–2700 m . In British Columbia , the Sierra Nevada ( California ), and the Rocky Mountains ( Colorado ), adults have been collected from early July through mid-August. In coastal California , captures are from October and November. Etymology. The species name is a patronym for Barbara Bartell , collector of the holotype . Remarks. Based on the considerable differences in habitat, it is possible that low elevation, coastal populations of S. bartellae (i.e., California ) represent a different species than the high elevation, montane populations (i.e., Sierra Nevada and Rocky Mountains). However, because I can find no difference in facies, male secondary features, or genitalia, I am treating them all as conspecific.