Unexpected high species diversity among European stalked puffballs - a contribution to the phylogeny and taxonomy of the genus Tulostoma (Agaricales) Author Jeppson, Mikael Author Altes, Alberto Author Moreno, Gabriel Author Nilsson, R. Henrik Author Yolanda Loarce, Author Bustos, Alfredo de Author Larsson, Ellen text MycoKeys 2017 21 33 88 http://dx.doi.org/10.3897/mycokeys.21.12176 journal article http://dx.doi.org/10.3897/mycokeys.21.12176 1314-4049--33 Tulostoma lloydii Bres., in Petri, Ann. Mycol. 2(5): 423. 1904. Figures 2h, 11 Holotype . USA (S!). A characteristic species originally described from N. America. We have not sequenced the holotype, but its macro- and micromorphology (Figure 11a) are very similar to that of the collections here studied (Figure 2h). Habitat and distribution. Tulostoma lloydii is recorded from N. America and Europe ( Wright 1987 ). In Europe it has a Mediterranean distribution, having been reported from Italy and Spain. According to Wright (1987) it is a species of "forest soil, among humus and plant debris". The studied material from Italy was collected in a garden, but there are also findings from more xeric environments on calcareous soil and on littoral sand dunes in Spain. Other specimens examined. ITALY, Friuli-Venezia Giulia: Pordenone, Vivaro, in litter under Thuja orientalis in church yard, Oct. 2012, E. Campo, M. Lahti 201210 (GB)*. SPAIN, Cordoba : Priego de Cordoba , terreno calizo con encinas, Apr. 1988, J. Gomez (AH 11606)*; Madrid: Guadalix de la Sierra, entre briofitos , en suelo basico , 23 May 2003, A. Gonzalez , ARBA (AH 31156, GB)*. Figure 11. T. lloydii , holotype (S): a spore ornamentation under SEM. Scale bar: 1 µm .