Taxonomic Study of Japanese Ctenopelmatinae (Hymenoptera, Ichneumonidae), with Descriptions of 22 New Species Author Watanabe, Kyohei text Bulletin of the Kanagawa Prefectural Museum (Natural Science) 2024 2024-03-22 53 39 102 journal article 10.5281/zenodo.13237363 2189-6720 13237363 D4FC7074-ADC6-4310-B60E-12B7ECBD6E25 Leipula pulchra sp. nov. (New SJN: Noudai-maru-himebachi) ( Figs. 29A–E , 39H , 40U ) Type series . Holotype : JAPAN , KPM-NK 84905 , F, Kanagawa Pref. , Atsugi City , Funako , campus of Tokyo University of Agriculture , 22. IV. – 16. V. 2016 , Y. Kato & S. Koizumi leg. (MsT) . Paratypes : JAPAN , KPM-NK 84906 , F, same data of holotype ; KPM-NK 84907 , 84908 , 2 F, ditto, 6. V. – 7. VI. 2016 , Y. Kato & S. Koizumi leg. (MsT) ; KPM-NK 84909 , M, Kanagawa Pref. , Odawara City , Kamisoga , 29. IV. 2018 , T. Amano leg. ; KPM-NK 84910 , F, Gifu Pref. , Toki City , Dachi , 22. IV. – 22. VI. 2007 , S. Takemoto leg. (MsT). Description. Female ( n = 5). Body length 5.9–7.6 (HT: 6.3) mm, polished and covered with dense punctures and silver setae. Head 0.6 × as long as wide in dorsal view. Clypeus 2.4– 2.5 (HT: 2.4) × as broad as high, rather sparsely punctate, weakly convex in lateral view, lower margin rounded ( Fig. 40U ). Face 1.7–1.8 (HT: 1.8) × as broad as high, almost flat in lateral view. Maximum length of gena in lateral view 0.6–0.7 (HT: 0.6) × basal width of mandible. Frons with a pair of conical tubercles ( Fig. 39H ). Dorsal profile of gena as Fig. 39H . Occipital carina complete. Length of malar space 0.6–0.8 (HT: 0.8) × as long as basal width of mandible. POL 1.0–1.2 (HT: 1.15) × as long as OD. OOL 1.9–2.0 (HT: 2.0) × as long as OD. Antenna with 37–40 (HT: 40) flagellomeres. FL I 6.65 × as long as maximum depth, 1.65–1.8 (HT: 1.65) × as long as FL II. Mesosoma. Epomia absent. Mesoscutum without notaulus. Epicnemial carina present laterally and ventrally, its dorsal end situated near anterior margin of mesopleuron. Speculum with large smooth area. Mesopleuron with irregular rugae around smooth speculum. Propodeum with no carinae except for posterior parts of lateromedian longitudinal carina ( Fig. 29C ). Pleural carina largely absent posteriorly. Fore wing length 4.8–5.8 (HT: 5.4) mm. Areolet present, shortly petiolated anteriorly, received vein 2m-cu slightly based of the outer angle ( Fig. 29D ). Fore wing vein 1cu-a interstitial or postfurcal (HT: postfurcal) to vein M&RS. Nervellus subvertical, intercepted at or slightly posterior the middle. Hind femur 4.9–5.2 (HT: 5.0) × as long as maximum depth in lateral view. Ratio of length of hind first to fifth tarsomeres 4.1–4.5 (HT: 4.1): 2.0: 1.3–1.4 (HT: 1.4): 0.8–0.9 (HT: 0.8): 1.2–1.35 (HT: 1.35). Fig. 29. Leipula pulchra sp. nov . (A–D: KPM-NK 84905, holotype, female; E: KPM-NK 84909, male) ― A: lateral habitus; B, E: head, frontal view; C: propodeum, dorso-lateral view; D: areolet. Metasoma. T I 2.35–2.75 (HT: 2.35) × as long as maximum width. T II 0.8–1.0 (HT: 0.8) × as long as maximum width. T III to T VII finely punctate. Ovipositor sheath 0.2–0.25 (HT: 0.25) × as long as hind tibia. Ovipositor with wide dorsal concavity. Colouration ( Figs. 29A–E ). Body (excluding wings and legs) black. Apex of mandibular teeth and ventral surfaces of scape and pedicel brown. Flagellum, posterior margin of T II, TIII except for small black area(s), membranous part of metasomal sternites, ovipositor sheath, and ovipositor yellowish brown to whitish yellow. Tegula whitish yellow. Sclerotized part of metasomal sternites sometimes partly tinged with dark brown. Wings hyaline. Veins and pterostigma blackish brown except for yellowish-brown wing base. Fore and mid legs reddish yellow except for coxae, trochanters and trochantelli partly to entirely yellow. Hind coxa reddish yellow, its apex more or less tinged with whitish yellow. Hind trochanter and trochantellus whitish yellow. Hind femur black except for narrow whitish yellow base. Hind tibia black except for subbasal whitish yellow band. Hind tibial spurs whitish yellow. Hind tarsus blackish brown to black, base of each segment narrowly tinged with whitish yellow. Whitish yellow areas of hind tarsus larger than female. Male ( n = 1). Similar to female. Face 1.6 × as broad as high. Length of malar space 0.5 × as long as basal width of mandible. OOL 1.7 × as long as OD. FL I 5.7 × as long as maximum depth. Hind femur 4.7 × as long as maximum depth in lateral view. Face, clypeus, and mandible except for teeth yellow ( Fig. 29E ). Distribution. Japan (Honshu). Bionomics. Unknown. Etymology . The specific name is from Latin “ pulchra ” (pretty and beautiful). Remarks. This species can be distinguished from other species by the frons with a pair of tubercles (absent in other species) and the T I 2.35–2.75 × as long as maximum width (other species shorter than 1.7 ×; see above key).