Taxonomic Study of Japanese Ctenopelmatinae (Hymenoptera, Ichneumonidae), with Descriptions of 22 New Species Author Watanabe, Kyohei text Bulletin of the Kanagawa Prefectural Museum (Natural Science) 2024 2024-03-22 53 39 102 journal article 10.5281/zenodo.13237363 2189-6720 13237363 D4FC7074-ADC6-4310-B60E-12B7ECBD6E25 Anisotacrus nocturnus sp. nov. (SJN: Kurayami-maru-himebachi) ( Figs. 6A–E , 38C , 40C ) Type series . Holotype : JAPAN , KPM-NK 84832 , F, Yamanashi Pref. , Sutama Town , Kanayamadaira , 25. VI. 2008 , T. Kidokoro leg. (LT) . Paratypes : JAPAN , KPM-NK 81350 , F, same data of holotype ; KPM-NK 84833 , F, ditto, 28. VI. 2008 , C. Satoh leg. (LT) . Description. Female ( n = 3). Body length 8.2–8.8 (HT: 8.2) mm, polished and covered with silver setae. Head 0.6 × as long as wide in dorsal view, finely coriaceous. Clypeus 2.35–2.5 (HT: 2.4) × as broad as high, sparsely punctate and polished, lower margin subtruncate medially ( Fig. 40C ). Face 1.75–1.8 (HT: 1.8) × as broad as high, weakly convex medially in lateral view. Frons with a median sharrow groove in front of median ocellus. Dorsal profile of gena as Fig. 38C . Occipital carina complete. Length of malar space 0.25–0.3 (HT: 0.3) × as long as basal width of mandible. Ocelli large ( Fig. 38C ). POL 0.7–0.75 (HT: 0.7) × as long as OD. OOL 0.45–0.5 (HT: 0.5) × as long as OD. Antenna with 34–36 (HT: 34) flagellomeres. FL I 5.7× as long as maximum depth, 1.65 × as long as FL II. Fig. 6. Anisotacrus nocturnus sp. nov. (KPM-NK 84832, holotype, female) ― A: lateral habitus; B: head, frontal view; C: mesopleuron; D: propodeum, dorso-lateral view; E: areolet. Mesosoma finely punctate and polished. Epomia absent. Mesoscutum with weak notaulus. Epicnemial carina present laterally and ventrally, its dorsal end situated near anterior margin of mesopleuron. Speculum with large smooth area ( Fig. 6C ). Propodeum finely punctate and coriaceous. Lateromedian longitudinal carina and lateral longitudinal carina absent anteriorly ( Fig. 6D ). Anterior transverse carina absent. Posterior transverse carina narrowly absent medially. Pleural carina weak. Area petiolaris smooth and 0.5 × as long as dorsal length of propodeum. Fore wing length 7.5–7.6 (HT: 7.6) mm. Areolet present, shortly petiolated anteriorly, received vein 2m-cu slightly based of the outer angle ( Fig. 6E ). Fore wing vein 1cu-a interstitial to vein M&RS ( Fig. 6A ). Nervellus inclivous, intercepted posterior the middle. Tarsal claws simple. Hind femur 5.6–6.35 (HT: 6.0) × as long as maximum depth in lateral view. Ratio of length of hind first to fifth tarsomeres 4.0–4.3 (HT: 4.1): 2.0: 1.45– 1.6 (HT: 1.4): 0.9–1.0 (HT: 0.95): 0.95–1.0 (HT: 1.0). Metasoma coriaceous. T I 1.9–2.0 (HT: 2.0) × as long as maximum width. T II 0.85–0.95 (HT: 0.95) × as long as maximum width. Ovipositor sheath 0.175–0.2 (HT: 0.2) × as long as hind tibia. Colouration ( Figs. 6A–E ). Body (excluding wings and legs) black. A pair of markings along frontal orbit and facial orbit, malar space, clypeus, mandible except for teeth, postero-dorsal and postero-ventral corners of pronotum, a pair of large markings of antero-lateral part of mesoscutum, tegula, and subtegular ridge yellow. Posterior margins of metasomal tergites narrowly tinged with yellow. Posterior corners of T III to T VI with whitish yellow marking. Membranous part of metasomal sternites, whitish yellow. Face sometimes with a pair of short longitudinal yellow stripes below antennal sockets. Wings hyaline. Veins and pterostigma blackish brown except for yellowish-brown wing base. Coxae and hind trochanter black (apices of fore and mid coxae usually with small yellow area). Fore and mid trochanters and trochantelli yellow. Fore and mid femora, tibiae, tibial spurs, and tarsi reddish yellow. Hind tibia and tibial spurs whitish yellow except for apical part of the former blackish brown. Hind tarsus blackish brown to black. Male. Unknown. Distribution. Japan (Honshu). Bionomics. Host is unknown. This species has large ocelli and collected by right trap. Thus, this species may be active in night. Etymology . The specific name is from Latin “ nocturnus ”. This species has nocturnal habit. Remarks. This species can be easily distinguished from other species by the large ocelli and body colouration.