Review of the genus Tylopus Jeekel, 1968, with descriptions of five new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae) Author Likhitrakarn, Natdanai Author Golovatch, Sergei I. Author Prateepasen, Rujiporn Author Panha, Somsak text ZooKeys 2010 72 23 68 http://dx.doi.org/10.3897/zookeys.72.744 journal article http://dx.doi.org/10.3897/zookeys.72.744 1313-2970-72-23 Tylopus parajeekeli sp. n. Figs 1315 Holotype ♂ (CUMZ), Thailand, Chiang Mai Province, Chom Thong District, Doi Inthanon National Park, summit, 2520 m, 18°34'29N, 98°28'48E, 12.10.2009, leg. S. Panha, J. Sutcharit & N. Likhitrakarn. Paratype: 1 ♂ (CUMZ), same locality, together with holotype. Name: To emphasize the close resemblance to Tylopus jeekeli Golovatch & Enghoff, 1993. Diagnosis: Very similar to Tylopus jeekeli , especially as regards its gonopod conformation, but differs in the paraterga lying much lower (at ca 1/3 versus 1/4-1/5 midbody height), in the caudal corners of the paraterga protruding behind the rear tergal contour already from segment 16 (versus segment 2), and also in gonopod spine z being much smaller and placed closer to the base of spine h. Description: Length 31 mm (holotype) or 31.5 mm (♂), width of midbody pro- and metazona 2.4 and 3.2 mm (holotype) or 2.3 and 3.4 mm (♂), respectively. Coloration of live animals and alcohol material uniformly blackish-brown (Fig. 13 A-G ); calluses of paraterga a little lighter, brown; antennomeres 1-5 light brown to yellowish, legs and venter light brown to grey-yellowish (Fig. 13 A-G ). All characters as in Tylopus bispinosus sp. n., except as follows. Antennae rather short and slender, reaching behind segment 3 dorsally. In width, head = segment 3 <collum <segments 2 and 4 <5-16; thereafter body gradually and gently tapering towards telson (Fig. 1B). Paraterga on collum like large rounded flaps (Fig. 13B, C). Following paraterga lying at about 1/3 midbody height, evidently declined ventrolaterally, subhorizontal only on a few posteriormost segments, mostly pointed caudally, subrectangular until segment 15, thereafter increasingly well protruding behind rear tergal contour (Fig. 13 A-G ). Metaterga with 2(3)+2(3) and 3 -5+3- 5 long setae arranged in two transverse rows. Axial line present on both halves of metaterga. Transverse sulcus present on segments 5-18, very finely beaded at bottom (Fig. 13B, F, G). Stricture between pro- and metazona finely striolate (Fig. 13B, D). Epiproct tip broad and emarginate (Fig. 13G, H). Hypoproct semi-circular, both caudal setae widely separated (Fig. 13H). Pleurosternal carinae as complete ridges on segments 2-4, thereafter retained until segment 17 mostly as a small caudal tooth (Fig. 13 C-F ). A low, only slightly divided, setose lobe between coxae 4 (Fig. 13I, J). Legs relatively short, ca 1.6-1.7 times as long as midbody height (Fig. 13K). Femora evidently bulged laterally (Fig. 13K); all postgonopodal legs except two last pairs with an evident adenostyle in parabasal 1/3 of each postfemur and tibia; tarsal brushes missing; all telopoditomeres except tarsi with dense adpressed pilosity (Fig. 13K). Gonopods (Figs 14, 15) with lobe l well-demarcated; spine h small, but elongate, not bifid; spine z very small, dentiform, placed at base of spine h. Figure 13. Tylopus parajeekeli sp. n., ♂ holotype (A) and ♂ paratype ( B-K ). A habitus, live coloration B, C anterior part of body, dorsal and lateral views, respectively D, E segments 10 and 11, dorsal and lateral views, respectively F, G, H posterior part of body, lateral, dorsal and ventral views, respectively I, J sternal cones between coxae 4, subcaudal and sublateral views, respectively K midbody leg. Figure 14. Tylopus parajeekeli sp. n., ♂ paratype. A, B right gonopod, mesal and lateral views, respectively C-F distal part of right gonopod, submesal, sublateral, subcaudal and suboral views, respectively. Scale bar: 0.2 mm. Figure 15. Tylopus parajeekeli sp. n., ♂ paratype. A, B right gonopod, lateral and mesal views, respectively. Scale bar: 0.5 mm.