South American Coccinellidae (Coleoptera), Part XIX: Overview of Cryptognathini and systematic revision of South American Cryptognatha Mulsant Author F, Guillermo González Author Hanley, Guy A. Author Gordon, Robert D. text Insecta Mundi 2019 2019-06-28 714 714 1 32 journal article 10.5281/zenodo.3674855 ac53680f-6358-4d6b-8880-ccf3dd73297e 1942-1354 3674855 F9BABD1C-B18B-47FD-9B87-F44C49FD9D30 Key to Cryptognatha species 1. Elytron immaculate, entirely yellow, brownish yellow, or black......................... 2 — Elytron maculate, with pale maculae on dark background, or with dark maculae on pale back- ground................................................................... 3 2(1). Elytron entirely yellow, brownish yellow ( Fig. 1 )............... 1. C. auriculata Mulsant — Elytron entirely black.......................................................... 13 3(1). Elytron with pale maculae on dark background...................................... 9 — Elytron with dark maculae on pale background...................................... 4 4(3). Maculae simple, usually single macula on lateral margin, occasionally with indistinct macula on sutural basal border...................................................... 5 — Maculae composed of multiple maculae on elytral surface............................. 7 5(4). Maculae consisting of short, dark maculae on lateral border outside of humeral callus and sometimes with indistinct sutural macula in basal 1/2 ( Fig. 55 )................................................................. 10. C. karla González and Hanley , n. sp. — Maculae consisting of single, complete macula on or near lateral margin from base of elytron to sutural margin in apical 1/2................................................... 6 6(5). Lateral macula wide, reaching lateral margin ( Fig. 26 )............. 5. C. amicta Gorham — Lateral macula separated from lateral margin ( Fig. 21 )........................................................................ 4. C. pam González and Hanley , n. sp. 7(4). Maculae consisting of two short vittae in basal 1/2 of elytron, single macula in apical 1/2, and irregular sutural macula ( Fig. 15 )......................... 3. C. gemellata Mulsant — Maculae differently arranged.................................................... 8 8(7). Dorsal pattern consisting of single large macula on basal ½ of elytron, two small maculae in basal apical 1/2, and irregular sutural vitta ( Fig. 110 )..................................................................... 21. C. vicki González and Hanley , n. sp. — Dorsal pattern consisting of irregularly arranged vittae with three more or less fused vittae in basal 1/2 becoming an irregular, large macula in basal 1/2 ( Fig. 8 ).......................................................................... 2. C. nodiceps Marshall 9(3). Elytron mostly dark with lateral margin pale...................................... 10 — Elytron with distinct central color pattern......................................... 12 10(9). Pale lateral margin of elytron wide, reddish yellow; pronotum entirely yellowish red ( Fig. 106 )....................................... 20. C. della González and Hanley, n. sp. — Pale lateral margin of elytron narrow, yellow; pronotum yellow with basomedian black macula................................................................... 11 11(10). Lateral margin on elytron sharply defined ( Fig. 40 ); Peru ................................................................... 7. C. hannah González and Hanley , n. sp. — Lateral margin on elytron obscure defined ( Fig. 32 ); Guyana ................................................................... 6. C. kellie González and Hanley , n. sp. 12(9). Elytron with basal 1/2 dark brown, apical 1/2 yellow (Fig. 49)........... 9. C. reedi Crotch — Elytron with elongate yellow macula next to scutellar shield, macula narrowly extended posteriorly parallel to suture ending in triangular macula at base of apical declivity ( Fig. 103 ).................................. 19. C. gayle González and Hanley , n. sp. 13(2). Pronotum entirely or mostly yellow.............................................. 14 — Pronotum mostly dark brown or black............................................ 15 14(13). Pronotum entirely yellow ( Fig. 99 ); Brazil ......................... 18. C. batesi Crotch — Pronotum mostly yellow with small, apically sinuate basal macula (Fig. 44).................................................... 8. C. whitney González and Hanley , n. sp. 15(13). Length 3.4 mm ; Colombia ( Fig. 113 )........................... 22. C. aethiops Crotch — Length less than 3.4 mm ; Brazil ................................................. 16 16(15). Male pronotum with narrow, yellow anterior border ( Fig. 82 )................................................................ 15. C. shelia González and Hanley , n. sp. — Male pronotum entirely black or dark brown....................................... 17 17(16). Penis guide of male genitalia abruptly narrowed in apical 1/2 ( Fig. 75 )................. 18 — Penis guide of male genitalia parallel-sided for most of length, or gradually narrowed in apical 1/2 ( Fig. 70 )............................................................... 20 18(17). Penis guide of male genitalia much shorter than paramere, apex very acute ( Fig. 75, 76 )........................................................... 13. C. flaviceps Crotch — Penis guide of male genitalia about at long as paramere............................. 19 19(18). Penis guide with apex rounded ( Fig. 79 ), weakly curved downward in lateral view ( Fig. 80 )..................................................... 14. C. weisei Brethes — Penis guide of male genitalia abruptly narrowed ( Fig. 64 ), sinuate in lateral view ( Fig. 64 )............................................................ 11. C. clarki Crotch 20(17). Penis guide of male genitalia about as long as paramere, oval, slightly widened in apical 1/2, then narrowed to weakly emarginate apex ( Fig. 70 )....................................................................... 12. C. celia González and Hanley , n. sp. — Penis guide of male genitalia long slender......................................... 21 21(20). Male genitalia with phallobase long, penis guide slightly longer than paramere, slender, evenly narrowed from base to abruptly rounded apex ( Fig. 88 )................................................................. 16. C. yolandi González and Hanley , n. sp. — Male genitalia elongate, penis guide about as long as paramere, oval, sides parallel in basal 1/2, curved to rounded apex in apical 1/2 ( Fig. 95 )........................... 17. C. pudibunda Mulsant