A revision of the Thyropygus allevatus group. Part 1: the T. opinatus subgroup (Diplopoda: Spirostreptida: Harpagophoridae)
Author
Pimvichai, Piyatida
Author
Enghoff, Henrik
Author
Panha, Somsak
text
Zootaxa
2009
2016
17
50
journal article
10.5281/zenodo.185971
1db39382-b4f0-46fa-ae08-dd5e57e0e26b
1175-5326
185971
Thyropygus implicatus
(
Demange, 1961
)
,
n. comb.
(
Figs. 2
,
3
,
8
A–D)
Cornugonus implicatus
Demange, 1961
: 181
.
Holotype
(not seen) in the Natural History Museum, London.
Material:
10 males
,
5 females
, and
1 juvenile
THAILAND
, Satun Province, Thale Ban National Park, 6° 42ˏ 29˝ N, 100° 10ˏ 46˝ E.
15 October 1991
. M. Andersen, O. Martin and N. Scharff leg., (
ZMUC
).
Diagnosis:
A species of the
opinatus
subgroup. Spatulate lobe (
sl
) at the apical part of telopodite terminating in a sharp brown spine. Similar in this respect to
T. opinatus
,
T. floweri
and
T. erectus
. Differs from these species by the having the lateral process of the anterior coxal fold (
alp
) apically bent abruptly mesad, its tip with serrate margins.
Description:
Adult males with 63–70 podous rings, no apodous rings. Length ca.
12–13 cm
, width ca. 7.4–8.0 mm. Adult females with 65–80 podous rings, no apodous rings. Length ca.
12–22 cm
, width ca. 8.0–11.0 mm. Color in life unknown; preserved specimens with head, antennae, legs, prozona, middorsal metazona, epiproct, paraprocts and hyproproct brown; metazona, margins of epiproct and paraprocts, and the tip of hypoproct darker brown.
Gonopods (
Figs. 8
A–D): Anterior coxal fold (
ac
) (
Fig. 8
A): lateral process (
alp
) apically bent abruptly mesad, tip with serrate margins, posterior side with a horizontal crest; mesal process (
amp
) much shorter than lateral process, directed meso-distad, pointed. Posterior coxal fold (
pc
) (
Fig. 8
B) with lateral paracoxites (
px
), digitiform; mesal process (
pmp
) complex, strongly developed along an anterior-posterior axis, with a large posterior broad lobe (
pbl
) and an anterior slender digitiform process (
adp
); lateral process (
plp
) small, pointed. Telopodite (
Figs. 8
C–D) leaving coxite between
pmp
and
plp
; femoral spine (
fe
) basally massive, ending in sharp point, directed almost straightly mesad,
in situ
resting against posterior surface of
alp
, fitting under its horizontal crest, crossing over with
fe
of the opposite side; telopodite distally to
fe
with a large, round lobe (
lo
) projecting distolaterally; tibial spine (
ti
) long, slender, curving in horizontal plane, its tip upturned and resting against anterior surface of
adp
; apical part: spatulate lobe (
sl
) terminating in a sharp brown spine; palette (
pa
) simple, gutter-like, distally with about nine brownish blepharochaetae (
bp
).
Distribution
(
Fig. 20
): Described from
MALAYSIA
, Penang Hill (5° 25ˏ 29˝ N, 100° 16ˏ 7˝ E) (
Demange, 1961
), based on specimens (including
holotype
) belonging to the Natural History Museum in London, and further specimens from
MALAYSIA
,
Malacca
, Sultanate of Kedah (2° 27ˏ 16˝ N, 102° 25ˏ 47˝ E) belonging to Naturalis, Leiden. The species was recorded as new for the
Thai
fauna by
Enghoff (2005)
based on the specimens studied by us (
THAILAND
, Satun Province, Thale Ban National Park).