Ranunculus luanchuanensis (Ranunculaceae), a new species from Henan, China
Author
Fei, Wen-Qun
Key Laboratory of Plant Resources Conservation and Sustainable Utilization, South China Botanical Garden, Chinese Academy of Sciences, Guangzhou 510655, Guangdong, China & University of Chinese Academy of Sciences, Beijing 100049, China
Author
Yuan, Qiong
https://orcid.org/0000-0003-2613-5045
Key Laboratory of Plant Resources Conservation and Sustainable Utilization, South China Botanical Garden, Chinese Academy of Sciences, Guangzhou 510655, Guangdong, China & Center of Conservation Biology, Core Botanical Gardens, South China Botanical Garden, Chinese Academy of Sciences, Guangzhou 510655, Guangdong, China
yuanqiong@scbg.ac.cn
Author
Yang, Qin-Er
https://orcid.org/0000-0002-6261-0731
Key Laboratory of Plant Resources Conservation and Sustainable Utilization, South China Botanical Garden, Chinese Academy of Sciences, Guangzhou 510655, Guangdong, China & Center of Conservation Biology, Core Botanical Gardens, South China Botanical Garden, Chinese Academy of Sciences, Guangzhou 510655, Guangdong, China
text
PhytoKeys
2023
2023-02-22
220
17
29
http://dx.doi.org/10.3897/phytokeys.220.96799
journal article
http://dx.doi.org/10.3897/phytokeys.220.96799
1314-2003-220-17
AF7CAAFAD50156C9AFCE52B668E110D4
Ranunculus luanchuanensis W.Q.Fei, Q.Yuan & Q.E.Yang
sp. nov.
Figs 1
, 2
, 3
, 4
Diagnosis.
The new species is morphologically similar to
R. limprichtii
in having 3-lobed and subreniform basal leaves, 3-lobed cauline leaves, and small flowers with reflexed and caducous sepals, but differs by having slender and basally slightly thickened roots (vs. fusiform), prostrate stems (vs. erect), obliquely ovoid and glabrous carpels and achenes (vs. widely ovoid and puberulous), longer styles in the carpels (ca. 1.2 mm vs. 0.6-0.8 mm) and achenes (ca. 1.8 mm vs. 0.6-0.8 mm), and glabrous receptacles (vs. sparsely puberulous).
Type
.
China
.
Henan province
:
Luanchuan county
,
Laojun Shan
,
33°43'3.62"N
,
111°38'59.89"E
, in shady place among boulders on mountaintop, alt.
2077 m
,
17 June 2022
,
W.Q. Fei
588
(
holotype
: IBSC; isotypes: IBSC, PE)
.
Description.
Herbs
perennial, terrestrial or rupicolous.
Roots
1-3, fibrous, slender, slightly thickened at base.
Stems
12-20 cm long, prostrate, unbranched, glabrous.
Basal leaves
2-6, 3-lobed, long petiolate; petioles 2-6 cm long, glabrous; blades 0.7-1.6
x
1-2.3 cm, subreniform in outline, thinly papery, adaxially green, glabrous or sparsely puberulous, abaxially light green, glabrous, base cordate, central segment 0.5-0.6
x
0.6-0.7 cm, widely obovate to rhombic-obovate, entire or 2-3-dentate, apex rounded or acuminate, lateral segments 0.5-0.7
x
0.7-1 cm, obliquely flabellate, inconspicuously 2-lobed, apex rounded or acuminate.
Lower cauline leaves
2-3, similar to basal ones but smaller.
Upper cauline leaves
1-2, 0.6-1.2
x
0.3-0.8 cm, 3-lobed, rarely entire, shortly petiolate or subsessile, oblate-ovate, flabellate or lanceolate, glabrous.
Inflorescences
terminal, 1(-2)-flowered.
Flowers
6-7 mm in diameter; pedicels 0.5-2 cm long, glabrous or sparsely puberulous; receptacles ca. 1 mm long, clavate, glabrous; sepals 5, 2-2.5
x
1.5-1.8 mm, elliptic to obovate, reflexed, caducous, green tinged with yellowish, concave, adaxially glabrous, abaxially sparsely puberulous; petals 5(-6), 3-3.5
x
1.5-1.8 mm, narrowly obovate, yellow, glabrous, apex obtuse or acuminate, nectary pit without a scale, claws ca. 0.4 mm long; stamens 12-16, filaments ca. 1.5 mm long, narrowly linear, anthers ca. 0.5 mm long, oblong; gynoecium subglobose; carpels 14-18, ovaries ca. 0.8 mm long, obliquely ovoid, laterally flattened, biconvex, glabrous, styles ca. 1.2 mm long, glabrous, apex recurved.
Aggregate fruit
ca. 3.5
x
3.5 mm, subglobose; achenes ca. 1.8
x
1.2 mm, obliquely ovoid, laterally flattened, biconvex, glabrous, styles ca. 1.8 mm long, persistent, apex recurved.
Etymology.
The specific epithet refers to the type locality of the new species, i.e., Luanchuan county in Henan province, central China.
Phenology.
Flowering in early June; fruiting at the end of June.
Distribution and habitat.
Ranunculus luanchuanensis
is currently known only from its type locality, i.e., Laojun Shan in Luanchuan county, Henan province, central China (Fig.
9
). It grows in a shady area among boulders on a mountaintop at an altitude of 2077 m above sea level.
Figure 9.
Distribution of
Ranunculus limprichtii
(black circle) and
R. luanchuanensis
sp. nov. (black square). Black arrow indicates the type locality of
R. limprichtii
, i.e., Dawu in Sichuan province, China.
Conservation status.
Ranunculus luanchuanensis
is currently known only from one small population at its type locality, i.e., Laojun Shan in Luanchuan county, Henan province, central China. This population consists of ca. 100 individuals within an area of less than 3 m2. However, the threat risk seems low because this species is not economically valuable and grows in a secluded place. The conservation status of
R. luanchuanensis
is here categorized as "Data Deficient (DD)" before adequate information on this species is acquired (
IUCN Standards and Petitions Committee 2022
).
Notes.
Since its description,
Ranunculus limprichtii var. flavus
Hand.-Mazz. has been known only from its type material from Songpan county in Sichuan province, China (
Wang 1995b
). Based on our observations of herbarium specimens and living plants in the wild, we agree with
Liou (1980)
that this variety should be reduced to the synonymy of
R. limprichtii
. We will deal with the identity of
R. limprichtii var. flavus
in detail elsewhere.
According to
Tamura's
(1995)
infrageneric classification of
Ranunculus
,
R. luanchuanensis
should be assigned to
R. sect. Ranunculus
, which is characterized by having swollen achenes with a distinct beak and receptacles hardly enlarged after anthesis.
Ranunculus limprichtii
, the putative closest ally of
R. luanchuanensis
, was placed by
Wang (1995b)
in
R. sect. Ranunculus
, with the section being incorrectly treated by him as
R. sect. Auricomus
(Spach) Schur. We accept the sectional placement of
R. limprichtii
since it is in accordance with the current placement of our new species in the same section.