Three new species of Kynotus from the Central Highlands of Madagascar (Clitellata, Megadrili) Author Csuzdi, Csaba 86F75D9D-740D-4D98-849B-B744C4438D76 Department of Zoology, Eszterházy Károly University, Eger, Hungary. & Email: csuzdi. csaba @ ektf. hu & urn: lsid: zoobank. org: author: 86 F 75 D 9 D- 740 D- 4 D 98 - 849 B-B 744 C 4438 D 76 csuzdi.csaba@ektf.hu Author Razafindrakoto, Malalatiana F21C5094-DE1F-4F7E-BF0A-C59C8AAF7D7A Department of Animal Biology, University of Antananarivo, Madagascar and Laboratory of Radio-Isotopes, University of Antananarivo, Madagascar. & Email: malalasraz @ yahoo. fr & urn: lsid: zoobank. org: author: F 21 C 5094 - DE 1 F- 4 F 7 E-BF 0 A-C 59 C 8 AAF 7 D 7 A malalasraz@yahoo.fr Author Hong, Yong 49F2D6FB-B6BA-438C-A452-ED4C109DB544 Department of Agricultural Biology, College of Agriculture & Life Sciences, Chonbuk National University, Jeonju, Republic of Korea. & urn: lsid: zoobank. org: author: 49 F 2 D 6 FB-B 6 BA- 438 C-A 452 - ED 4 C 109 DB 544 & Corresponding author: geoworm @ hanmail. net geoworm@hanmail.net text European Journal of Taxonomy 2017 2017-07-13 336 1 14 journal article 22086 10.5852/ejt.2017.336 4f3fdc50-52dc-4b53-9e5e-cbcc9b4e5985 2118-9773 3829873 A61DAF10-1819-4C90-A64A-1F1DDDAC220D Kynotus ankisiranus sp. nov. urn:lsid:zoobank.org:act: DDAFA680-A9E8-4A8B-AD38-9FB52F7378F2 Figs 2 , 5 Diagnosis Length 70–90 mm , diameter 3 mm . Colour grey with slight red pigmentation at the head on dorsum. Segments 1–3 simple, 4–10 biannulate. Setae appear from segment 2 and clearly visible, male pores ventral on 16. Clitellum circular on 18–27, ½28. Three rows of spermathecae of elongated oval-shape in 13/14, 14/15, 15/16. No septa notably thickened. Genital setal glands in 14, 15. Etymology The species epithet refers to the type locality, Mt Ankisira National Park. Table 1. Localities and COI Genbank accession numbers of the specimens of Kynotus analysed (EWSJC samples were obtained by Sam James).
Species Voucher no. Locality Accession no.
K. ankisiranus sp. nov. HNHM/AF5659 Madagascar, Ambalavao, Namoly, Mt Ankisira National Park KX527886
K. alaotranus 1 HNHM AF/5528 Madagascar, Vohimana Reserve, secondary Pandanus forest KX372247
K. alaotranus 2 HNHM AF/5528 Madagascar, Vohimana Reserve, secondary Pandanus forest KX372248
K. blancharti sp. nov. HNHM/AF5539 Madagascar, Antsirabe District, Sambaina KX517831
K. blancharti sp. nov. HNHM/AF5539 Madagascar, Antsirabe District, Sambaina KX517832
K. michaelseni 1 HNHM AF/5530 Madagascar, Sambaina, open bush-land KX372243
K. michaelseni 2 HNHM AF/5530 Madagascar, Sambaina, open bush-land KX372244
K. minutus 1 HNHM AF/5235 Madagascar, Vohimana Reserve, secondary Pandanus forest KX372245
K. minutus 2 HNHM AF/5235 Madagascar, Vohimana Reserve, secondary Pandanus forest KX372246
K. proboscideus EWSJC848 Madagascar, Mangalaza Ambatondrazaka, near villages JN260829
K. proboscideus EWSJC849 Madagascar, Mangalaza Ambatondrazaka, near villages JN260830
K. proboscideus EWSJC850 Madagascar, Mangalaza Ambatondrazaka, near villages JN260831
K. proboscideus EWSJC851 Madagascar, Mangalaza Ambatondrazaka, near villages JN260832
K. proboscideus EWSJC852 Madagascar, Mangalaza Ambatondrazaka, near villages JN260833
K. proboscideus EWSJC855 Madagascar, Vohitsoa Ambatondrazaka, along the road JN260834
K. proboscideus HNHM AF/5512 Madagascar, Vohitsoa Ambatondrazaka, Tsarahonenana KX527885
K. sihanakus EWSJC924 Madagascar, Bemanevika Bealanana, grassland JN260884
K. sihanakus EWSJC925 Madagascar, Bemanevika Bealanana, grassland JN260885
K. sihanakus EWSJC927 Madagascar, Bemanevika Bealanana, grassland JN260886
K. sihanakus EWSJC928 Madagascar, Bemanevika Bealanana, grassland JN260887
K. sihanakus EWSJC929 Madagascar, Bemanevika Bealanana, grassland JN260888
K. sihanakus EWSJC930 Madagascar, Ambatosoratra Ambatondrazaka, banana plantation JN260889
K. sihanakus EWSJC931 Madagascar, Ambatosoratra Ambatondrazaka, banana plantation JN260890
K. sihanakus EWSJC939 Madagascar, Ambatosoratra Ambatondrazaka, banana plantation JN260895
K. voimmanus sp. nov. HNHM AF/5661 Madagascar, Moramanga, Andasibe, Voimma Community Park KX527887
K. voimmanus sp. nov. HNHM AF/5661 Madagascar, Moramanga, Andasibe, Voimma Community Park KX517833
K. voimmanus sp. nov. HNHM AF/5661 Madagascar, Moramanga, Andasibe, Voimma Community Park KX517834
K. voimmanus sp. nov. HNHM AF/5661 Madagascar, Moramanga, Andasibe, Voimma Community Park KX517835
K. voimmanus sp. nov. HNHM AF/5662 Madagascar, Moramanga, Andasibe, Voimma Community Park KX517836
K. voimmanus sp. nov. HNHM AF/5662 Madagascar, Moramanga, Andasibe, Voimma Community Park KX517837
K. voimmanus sp. nov. HNHM AF/5662 Madagascar, Moramanga, Andasibe, Voimma Community Park KX517838
Material examined Holotype MADAGASCAR : clitellate, Ambalavao District , Sendreisoa , Namoly , Mt Ankisira National Park , Aristida grassland at the edge of primary forest, 22°06′45.8″ S , 46°56′33.8″ E , 1500 m a.s.l. , 24 Feb. 2015 , Y. Hong and M. Razafindrakoto leg. ( HNHM / AF5658 ). Paratypes MADAGASCAR : 2 clitellate, 3 aclitellate adults ( HNHM /AF5659), 2 aclitellate ( CHBNU MD- 06- AL5), locality and date same as for holotype . Description Holotype 50 mm in length, diameter after clitellum 3 mm , segment number 98, tail missing. Paratypes 72–86 mm in length, 3 mm in diameter, segment number 170–179. Colour grey with red pigmentation at head on dorsum. Head zygolobous, segments 1–3 simple, 4–10 biannulate. Dorsal pores lacking. Setae small, ab and cd clearly observed from segment 2. Setal arrangement ( holotype ) after clitellum aa:ab:bc:cd:dd = 15:1.25:9.5:1:18, dd = 0.32 U ( Fig. 2A ). Clitellum annular on segments 18–27, ½ 28. Fig. 2. Kynotus ankisiranus sp. nov. A . Setal arrangement, a,b,c, d representing setal lines. B .Spermathecae, schematic representation. C . Genital seta. D . Tip of the genital seta. Scale bars = 0.1 mm. Male pores ventral on 16. Female pores not seen. Spermathecal pores in 13/14 (2), 14/15 (2), 15/16 (2) from bc to cd . Genital setal pores in variable position between ab and cd on 14, 15. Internal characters Large muscular gizzard in 5. Septa all membranous. Calciferous glands, lamellae and typhlosolis lacking. Dorsal blood vessel simple throughout, last pair of hearts in 11. Excretory system holoic, vesiculate. Two pairs of testes and sperm funnels in 10, 11 enclosed in large peri-oesophageal testis sacs. Seminal vesicles lacking. Ovaries in 13. A pair of large, oval copulatory chambers occupying the ventral space of segments 16–17. Each copulatory chamber bearing an irregular prostate-like gland (pseudoprostate) reaching segment 21. Spermathecae elongated, oval-shaped, with short duct attached at 13/14(2), 14/15(2), 15/16(2) in variable position arranged from between bc to cd ( Fig. 2B ). Two pairs of genital setal glands present in segments 14, 15 above ab and around cd respectively. The genital setae are slightly curved, with spoon-shaped apex, length 1.6–1.7 mm , diameter at the middle 0.05 mm , ornamentation dense serrations ( Fig. 2 C–D). Remarks The new species belongs to the alaotranus group ( Table 2 ) characterized by biannulate segments between 4–10. With its smaller size and red pigmentation on the dorsum, K. ankisiranus sp. nov. most closely resembles K. minutus Csuzdi, Razafindrakoto & Blanchart, 2012 , but differs from it by the longer clitellum (18–26 vs 18–27, ½ 28) and the number of genital setal glands (single pair in 15 vs two pairs in 14, 15). K. amkisiranus sp. nov. is clearly different from all the examined alaotranus group’s species by its COI barcode as well ( Table 3 , Fig. 5 ).