Reassessing the phylogeny and divergence times of sloths (Mammalia: Pilosa: Folivora), exploring alternative morphological partitioning and dating models Author Casali, Daniel M Author Boscaini, Alberto Author Gaudin, Timothy J Author Perini, Fernando A text Zoological Journal of the Linnean Society 2022 2022-12-01 196 4 1505 1551 https://academic.oup.com/zoolinnean/article/196/4/1505/6617197 journal article 199613 10.1093/zoolinnean/zlac041 c62342a9-9254-4518-a373-588a3d66fb25 0024-4082 7381236 Megalonychidae PP = 63, age = 20.88 Mya (18.61–23.61). The position of Hapalops as a stem megalonychid was recovered in all BI analyses, but was poorly supported. The clade containing all other Megalonychidae will be detailed below. Megalonychidae was supported by seven synapomorphies (one for both methods and six exclusively for BI): Cf1 closer to anterior edge of maxilla than to Mf1 (BI); maximum height of coronoid process greater than its anteroposterior length (BI); large supraoccipital exposure on cranial roof, expanded anteriorly at midline; triangular occiput in posterior view (BI); presence of paroccipital process foramen (BI); eustachian tube opening formed by entotympanic and ectotympanic (BI); and absence of a deep notch for medial cruciate (posterior) ligament in femur (BI).