<document id="B0B862DD176346B0FB4F0004E902D4E3" ID-CLB-Dataset="7631" ID-DOI="10.1206/0003-0090-423.1.1" ID-GBIF-Dataset="2e886aea-b59a-45a6-aeaa-2427d584a894" ID-ISSN="0003-0090" ID-Zenodo-Dep="4613008" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615995240537" checkinUser="felipe" docAuthor="Grimaldi, David A." docDate="2018" docId="03B187A8FFB3FFC7FD7D3B1A7FB7FDF5" docLanguage="en" docName="BulAmeMusNatHis.2018.423.1-97.pdf.imf" docOrigin="Bulletin of the American Museum of Natural History 2018 (423)" docSource="http://www.bioone.org/doi/10.1206/0003-0090-423.1.1" docStyle="DocumentStyle:C5E2DA72A22EF33813C92A197453A310.5:BulAmeMusNatHis.2011-.journal_article.0cover" docStyleId="C5E2DA72A22EF33813C92A197453A310" docStyleName="BulAmeMusNatHis.2011-.journal_article.0cover" docStyleVersion="5" docTitle="Lindneromyia neomedialis Grimaldi 2018, new species" docType="treatment" docVersion="7" lastPageNumber="97" masterDocId="FF88FFD0FF86FFFEFFAA39537F49FFF7" masterDocTitle="Basal Cyclorrhapha In Amber From The Cretaceous And Tertiary (Insecta: Diptera), And Their Relationships: Brachycera In Cretaceous Amber Part Ix David A. Grimaldi" masterLastPageNumber="97" masterPageNumber="1" pageNumber="1" updateTime="1698933720305" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="72B2C39220CF8D9E3E7F271809541633">Basal Cyclorrhapha In Amber From The Cretaceous And Tertiary (Insecta: Diptera), And Their Relationships: Brachycera In Cretaceous Amber Part Ix David A. Grimaldi</mods:title>
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<mods:name id="74466EBAD643D8EDA9021A3F9D721B59" type="personal">
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<mods:namePart id="86F9392E0A1EE52F197C16E66F179166">Grimaldi, David A.</mods:namePart>
<mods:affiliation id="9E1DAF1941163229DFD23493C5B91D16">Division of Invertebrate Zoology American Museum of Natural History, New York</mods:affiliation>
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<treatment id="03B187A8FFB3FFC7FD7D3B1A7FB7FDF5" ID-DOI="http://doi.org/10.5281/zenodo.4631133" ID-GBIF-Taxon="180670801" ID-Zenodo-Dep="4631133" LSID="urn:lsid:plazi:treatment:03B187A8FFB3FFC7FD7D3B1A7FB7FDF5" httpUri="http://treatment.plazi.org/id/03B187A8FFB3FFC7FD7D3B1A7FB7FDF5" lastPageId="57" pageId="53">
<subSubSection id="C3026535FFB3FFCBFD7D3B1A7BD8FD95" box="[727,1169,585,610]" pageId="53" type="nomenclature">
<paragraph id="8BA736BEFFB3FFCBFD7D3B1A7BD8FD95" blockId="53.[727,1169,585,610]" box="[727,1169,585,610]" pageId="53">
<heading id="D0EF81D2FFB3FFCBFD7D3B1A7BD8FD95" box="[727,1169,585,610]" centered="true" fontSize="9" level="2" pageId="53" reason="2">
<taxonomicName id="4C184D3DFFB3FFCBFD7D3B1A7B4BFD96" ID-CoL="8CMH3" authority="Grimaldi, 2018" authorityName="Grimaldi" authorityYear="2018" box="[727,1026,585,609]" class="Insecta" family="Platypezidae" genus="Lindneromyia" kingdom="Animalia" order="Diptera" pageId="53" phylum="Arthropoda" rank="species" species="neomedialis" status="sp. nov.">
<emphasis id="B96CEAACFFB3FFCBFD7D3B1A7B4BFD96" bold="true" box="[727,1026,585,609]" italics="true" pageId="53">Lindneromyia neomedialis</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A25F57D7FFB3FFCBFBBA3B197BD8FD95" box="[1040,1169,586,610]" pageId="53" rank="species">new species</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3026535FFB3FFCBFCC43B287CB3FD63" box="[878,1018,635,660]" pageId="53" type="description">
<paragraph id="8BA736BEFFB3FFCBFCC43B287CB3FD63" blockId="53.[878,1018,635,660]" box="[878,1018,635,660]" pageId="53">
<heading id="D0EF81D2FFB3FFCBFCC43B287CB3FD63" box="[878,1018,635,660]" centered="true" fontSize="9" level="2" pageId="53" reason="2">
<figureCitation id="13232A3BFFB3FFCBFCC43B287C9DFD63" box="[878,980,635,660]" captionStart="FIG" captionStartId="16.[110,151,1649,1670]" captionTargetBox="[109,1213,226,1619]" captionTargetId="figure-9@16.[109,1213,226,1619]" captionTargetPageId="16" captionText="FIG. 9. Lindneromyia neomedialis, n. sp. (Platypezidae), male holotype in Miocene amber from the Dominican Republic, AMNH DR14-35. A. Dorsal habitus. B. Ventral habitus. C. Ventral view of head. D. Ventral view of terminalia and hind legs." figureDoi="http://doi.org/10.5281/zenodo.4613028" httpUri="https://zenodo.org/record/4613028/files/figure.png" pageId="53">Figures 9</figureCitation>
, 
<figureCitation id="13232A3BFFB3FFCBFC4A3B2F7CB3FD63" box="[992,1018,636,660]" captionStart="FIG" captionStartId="54.[107,149,1457,1478]" captionTargetBox="[191,1184,234,1426]" captionTargetId="figure-9@54.[108,1212,224,1431]" captionTargetPageId="54" captionText="FIG. 34. Lindneromyia neomedialis, n. sp. (Platypezidae), male holotype in Miocene amber from the Dominican Republic, AMNH DR14-35. A. Head, frontal view. B. Tip of palp. C, D, E. Tarsi: hind, mid, and fore (respectively). F. Apex of abdomen, ventral. G. Wing (slightly reconstructed at base)." figureDoi="http://doi.org/10.5281/zenodo.4613094" httpUri="https://zenodo.org/record/4613094/files/figure.png" pageId="53">34</figureCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3026535FFB3FFCBFD663BFE7BEAFBE6" pageId="53" type="diagnosis">
<paragraph id="8BA736BEFFB3FFCBFD663BFE7BEAFBE6" blockId="53.[684,1214,685,1738]" pageId="53">
DIAGNOSIS: (male only). Vein M unforked (M 
<subScript id="179C34FBFFB3FFCBFD643B897D9EFD1C" attach="left" box="[718,727,730,747]" fontSize="7" pageId="53">2</subScript>
lost), R 
<subScript id="179C34FBFFB3FFCBFC923B897C1DFD1C" attach="left" box="[824,852,730,747]" fontSize="7" pageId="53">4+5</subScript>
and M 
<subScript id="179C34FBFFB3FFCBFC043B897CFEFD1C" attach="left" box="[942,951,730,747]" fontSize="7" pageId="53">1</subScript>
slightly sinuous; gena completely and occiput virtually bare; metatarsomeres 1–4 flattened, expanded, basitarsomeres short, tarsomere 3 longest; acrostichals completely absent; only 1 (short) pair dorsocentrals and scutellars present; 2 notopleurals; wing membrane with very faint, minute microtrichia over about half of wing (mostly posteriorly); male abdominal segments 7–8 inserted into deep emargination of posterior portion of tergite 6.
</paragraph>
</subSubSection>
<subSubSection id="C3026535FFB3FFC9FD663D497BC4FD71" lastPageId="55" pageId="53" type="description">
<paragraph id="8BA736BEFFB3FFC9FD663D497E23FEEC" blockId="53.[684,1214,685,1738]" lastBlockId="55.[108,637,226,1738]" lastPageId="55" pageId="53">
DESCRIPTION: (male only). 
<emphasis id="B96CEAACFFB3FFCBFB8B3D497B2FFBC5" bold="true" box="[1057,1126,1050,1074]" pageId="53">Head:</emphasis>
Large, broader than thorax. Eyes bare, large, occupying all of lateral portion and much of frontal surface of head; holoptic in male from just above antennae to anterior ocellus (~15 facets); dorsal facets about 2× diameter of ventral ones. Median furrow of frons, interfrontal setae, orbitals, verticals (presence/absence) not observable. Ocelli on tubercle; setae absent, 6 minute setulae present. Postocular setae minute, in short row along posterodorsal margin of eye; occiput otherwise bare. Vibrissa, genal setae absent. 
<emphasis id="B96CEAACFFB3FFCBFC763CD47B0FFA68" bold="true" box="[988,1094,1415,1439]" pageId="53">Antenna:</emphasis>
Slight oval depression on face beneath each antenna. Scape small, bare; pedicel cup shaped, with 4–5 setulae on rim; arista terminal; basal flagellomere short, roughly drop shaped but without slender apical extension. Arista with two minute, ringlike articles at base; basal aristomere long, fine, bare (not micropubescent). 
<emphasis id="B96CEAACFFB3FFCBFCDB3F237CB6F97F" bold="true" box="[881,1023,1648,1672]" pageId="53">Mouthparts:</emphasis>
Oral margin and clypeus squared; clypeus thin. Palp small, with apex truncate (possibly with a pore). Labellum (opened) broad, with ~10 pseudotracheae; labrum not observable.
</paragraph>
<caption id="DF676636FFB0FFC8FFC13CE27C86F9F5" ID-DOI="http://doi.org/10.5281/zenodo.4613094" ID-Zenodo-Dep="4613094" httpUri="https://zenodo.org/record/4613094/files/figure.png" pageId="54" startId="54.[107,149,1457,1478]" targetBox="[191,1184,234,1426]" targetPageId="54">
<paragraph id="8BA736BEFFB0FFC8FFC13CE27C86F9F5" blockId="54.[107,1211,1455,1538]" pageId="54">
FIG. 34. 
<taxonomicName id="4C184D3DFFB0FFC8FF683CFC7E80FA30" authority="Grimaldi, 2018" authorityName="Grimaldi" authorityYear="2018" box="[194,457,1455,1479]" class="Insecta" family="Platypezidae" genus="Lindneromyia" kingdom="Animalia" order="Diptera" pageId="54" phylum="Arthropoda" rank="species" species="neomedialis" status="sp. nov.">
<emphasis id="B96CEAACFFB0FFC8FF683CFC7E80FA30" box="[194,457,1455,1479]" italics="true" pageId="54">Lindneromyia neomedialis</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A25F57D7FFB0FFC8FE7F3CFC7D42FA31" box="[469,523,1455,1478]" pageId="54" rank="species">n. sp.</taxonomicNameLabel>
(
<taxonomicName id="4C184D3DFFB0FFC8FDBD3CFC7DD4FA30" box="[535,669,1455,1479]" class="Insecta" family="Platypezidae" kingdom="Animalia" order="Diptera" pageId="54" phylum="Arthropoda" rank="family">Platypezidae</taxonomicName>
), male holotype in Miocene amber from the Dominican Republic, AMNH DR14-35. 
<emphasis id="B96CEAACFFB0FFC8FE6A3C9F7E90FA13" bold="true" box="[448,473,1484,1508]" pageId="54">A.</emphasis>
Head, frontal view. 
<emphasis id="B96CEAACFFB0FFC8FD053C9F7D8CFA13" bold="true" box="[687,709,1484,1508]" pageId="54">B.</emphasis>
Tip of palp. 
<emphasis id="B96CEAACFFB0FFC8FCFA3C9F7CEEFA13" bold="true" box="[848,935,1484,1508]" pageId="54">C, D, E.</emphasis>
Tarsi: hind, mid, and fore (respectively). 
<emphasis id="B96CEAACFFB0FFC8FEAA3CB97E47F9F5" bold="true" box="[256,270,1514,1538]" pageId="54">F</emphasis>
. Apex of abdomen, ventral. 
<emphasis id="B96CEAACFFB0FFC8FD9B3CBA7D0BF9F6" bold="true" box="[561,578,1513,1537]" pageId="54">G</emphasis>
. Wing (slightly reconstructed at base).
</paragraph>
</caption>
<paragraph id="8BA736BEFFB1FFC9FF2738777D35FB4F" blockId="55.[108,637,226,1738]" pageId="55">
<emphasis id="B96CEAACFFB1FFC9FF2738777FA1FECB" bold="true" box="[141,232,292,316]" pageId="55">Thorax:</emphasis>
Scutum rounded but not arched; curvature greater on anterior portion; scutum and scutellum appear black, perhaps velvety. Acrostichals absent; only one pair of (short, prescutellar, inclinate) dorsocentrals present (smaller graded dcs completely absent). Supraalar setae absent; 1 short postalar; 2 short notopleurals; anepisternum and katepisternum bare. Prescutellum absent. Scutellum with only one pair of small preapical setae, divergent. 
<emphasis id="B96CEAACFFB1FFC9FD9C3B1E7D35FD92" bold="true" box="[566,636,589,613]" pageId="55">Wing:</emphasis>
Length/width 2.0, hyaline except for faint tint between Sc and R 
<subScript id="179C34FBFFB1FFC9FE9E3BC87E74FD5B" attach="left" box="[308,317,667,684]" fontSize="7" pageId="55">1</subScript>
. Membrane with faint, minute, barely detectable microtrichia over about half the (posterior portion of) wing, including much of cell Sc. No veins with setulae or spinules; prehumeral costal seta very small; vein C with spinules to slightly past apex of R 
<subScript id="179C34FBFFB1FFC9FDB93A137D67FCA6" attach="left" box="[531,558,832,849]" fontSize="7" pageId="55">2+3</subScript>
; C terminates at apex of M 
<subScript id="179C34FBFFB1FFC9FEF33A327E2BFC85" attach="left" box="[345,354,865,882]" fontSize="7" pageId="55">1</subScript>
. Sc relatively short, close to R 
<subScript id="179C34FBFFB1FFC9FF373AD17FEFFC64" attach="left" box="[157,166,898,915]" fontSize="7" pageId="55">1</subScript>
; pterostigma absent. R 
<subScript id="179C34FBFFB1FFC9FE033AD17E8CFC64" attach="left" box="[425,453,898,915]" fontSize="7" pageId="55">2+3</subScript>
almost straight, R 
<subScript id="179C34FBFFB1FFC9FFD73AF07FD0FC43" attach="left" box="[125,153,931,948]" fontSize="7" pageId="55">4+5</subScript>
and M 
<subScript id="179C34FBFFB1FFC9FF4D3AF77FB9FC42" attach="left" box="[231,240,932,949]" fontSize="7" pageId="55">1</subScript>
slightly sinuous, convergent at wing tip; R 
<subScript id="179C34FBFFB1FFC9FF033A977F8CFC22" attach="left" box="[169,197,964,981]" fontSize="7" pageId="55">4+5</subScript>
ends near wing tip; cells br, bm closed. Crossvein dm-cu short, distant from wing margin, 5× its length from wing margin (vs. ca. 0.5–3×). M unforked (M 
<subScript id="179C34FBFFB1FFC9FE293D7B7EC5FBCE" attach="left" box="[387,396,1064,1081]" fontSize="7" pageId="55">2</subScript>
lost), without abrupt bend in M 
<subScript id="179C34FBFFB1FFC9FF433D1A7FBBFBAD" attach="left" box="[233,242,1097,1114]" fontSize="7" pageId="55">1</subScript>
. CuA 
<subScript id="179C34FBFFB1FFC9FE9D3D1A7E09FBAD" attach="left" box="[311,320,1097,1114]" fontSize="7" pageId="55">1</subScript>
relatively straight; cell cup pointed, CuP straight (not curved); anal lobe and alula well developed. Halter knob large, dark, bulbous, length ~ equal to that of stem.
</paragraph>
<paragraph id="8BA736BEFFB1FFC9FFC63D917D11F9F4" blockId="55.[108,637,226,1738]" pageId="55">
<emphasis id="B96CEAACFFB1FFC9FFC63D917FEEFB2D" bold="true" box="[108,167,1218,1242]" pageId="55">Legs:</emphasis>
Relatively short; all femora without macrosetae, spines or tubercles. Tarsomeres 1–4 flattened, expanded; tarsomere 3 longest, 1 is broadest. All tarsomeres with posteroventral, seamlike row of dark, bifid scales; more such scales scattered over dorsum of metatarsomeres; 2 sparse rows of scales dorsally on meso- and metatibia. Mesotibia with single, short ventroapical spur, length less than diameter of tibial apex. Pulvilli well developed; empodium setiform.
</paragraph>
<paragraph id="8BA736BEFFB1FFC9FF273F5F7B5BFE16" blockId="55.[108,637,226,1738]" lastBlockId="55.[684,1213,226,1738]" pageId="55">
<emphasis id="B96CEAACFFB1FFC9FF273F5F7E4DF9D3" bold="true" box="[141,260,1548,1572]" pageId="55">Abdomen:</emphasis>
Dark; relatively short, as preserved only slightly longer than thorax. Tergites well developed, wrap laterally, contact lateral margins of sternites (which are also well developed). Abdomen laterally with fine, erect setae; dorsally with few or no setae (probably velvety black); terminal tergites without large, spinelike setae. Six tergites visible dorsally, 2 small ones ventrally (tergites 7 and 8) (plus epandrium), all three ventroflexed (presumably completely circumverted), recessed in deep emargination on posterior portion of tergite 6. Cerci light, appear fused at bases. Posterior margin tergite 6 without thick, stiff setae; epandrium crescentic.
</paragraph>
<paragraph id="8BA736BEFFB1FFC9FD6738B97BC4FD71" blockId="55.[684,1213,226,1738]" pageId="55">
<materialsCitation id="3B703CE3FFB1FFC9FD6738B97BC4FD71" ID-GBIF-Occurrence="3059949302" collectionCode="AMNH" country="Dominican Republic" location="Specimen" pageId="55" specimenCode="DR14-35" specimenCount="1" stateProvince="Santiago" typeStatus="holotype">
SPECIMENS EXAMINED: 
<typeStatus id="54A3881CFFB1FFC9FC5238B97B2CFDF5" box="[1016,1125,490,514]" pageId="55" type="holotype">Holotype</typeStatus>
, male, 
<collectionCode id="ED09AE7BFFB1FFC9FD063B587C48FDD4" box="[684,769,523,547]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="55">AMNH</collectionCode>
<specimenCode id="DBBE9EC5FFB1FFC9FCA43B587C3DFDD4" box="[782,884,523,547]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="55">DR14-35</specimenCode>
, from mines near 
<collectingRegion id="49DCF85CFFB1FFC9FBFD3B587BF1FDD4" box="[1111,1208,523,547]" country="Dominican Republic" name="Santiago" pageId="55">Santiago</collectingRegion>
, 
<collectingCountry id="F30F762EFFB1FFC9FD063B7F7CD2FDB3" box="[684,923,556,580]" name="Dominican Republic" pageId="55">Dominican Republic</collectingCountry>
(Miocene). 
<location id="8EC76065FFB1FFC9FB9A3B7F7BD4FDB3" LSID="urn:lsid:plazi:treatment:03B187A8FFB3FFC7FD7D3B1A7FB7FDF5:8EC76065FFB1FFC9FB9A3B7F7BD4FDB3" box="[1072,1181,556,580]" country="Dominican Republic" name="Specimen" pageId="55" stateProvince="Santiago">Specimen</location>
is complete and has excellent preservation, although the surface of the amber is crazed.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3026535FFB1FFC9FD663BC37B62FCFC" pageId="55" type="etymology">
<paragraph id="8BA736BEFFB1FFC9FD663BC37B62FCFC" blockId="55.[684,1213,226,1738]" pageId="55">
ETYMOLOGY: 
<emphasis id="B96CEAACFFB1FFC9FCCF3BC37CC6FD50" box="[869,911,656,679]" italics="true" pageId="55">Neo</emphasis>
-, referring to New World; and - 
<emphasis id="B96CEAACFFB1FFC9FD403BE37C0DFD3F" box="[746,836,688,712]" italics="true" pageId="55">medialis</emphasis>
, referring to the medial vein, as this is the only New World species of the genus thus far with a simple medial vein.
</paragraph>
</subSubSection>
<subSubSection id="C3026535FFB1FFC7FD673A477FB7FDF5" lastPageId="57" pageId="55" type="discussion">
<paragraph id="8BA736BEFFB1FFC9FD673A477BF4F93D" blockId="55.[684,1213,226,1738]" pageId="55">
COMMENTS: 
<taxonomicName id="4C184D3DFFB1FFC9FCC23A477B4FFCDB" authorityName="Kessel" authorityYear="1965" box="[872,1030,788,812]" class="Insecta" family="Platypezidae" genus="Lindneromyia" kingdom="Animalia" order="Diptera" pageId="55" phylum="Arthropoda" rank="genus">
<emphasis id="B96CEAACFFB1FFC9FCC23A477B4FFCDB" box="[872,1030,788,812]" italics="true" pageId="55">Lindneromyia</emphasis>
</taxonomicName>
is the largest genus of the family, and the predominant one in the tropics. Three species in the genus have a simple medial vein: 
<taxonomicName id="4C184D3DFFB1FFC9FC343A2B7B26FC78" authority="Kessel" authorityName="Kessel" box="[926,1135,887,911]" class="Insecta" family="Platypezidae" genus="Lindneromyia" kingdom="Animalia" order="Diptera" pageId="55" phylum="Arthropoda" rank="species" species="africana">
<emphasis id="B96CEAACFFB1FFC9FC343A2B7CE5FC78" box="[926,940,888,911]" italics="true" pageId="55">L</emphasis>
. 
<emphasis id="B96CEAACFFB1FFC9FC6A3A247B55FC78" box="[960,1052,887,911]" italics="true" pageId="55">africana</emphasis>
Kessel
</taxonomicName>
(from 
<collectingCountry id="F30F762EFFB1FFC9FD063ACA7DBDFC46" box="[684,756,921,945]" name="Kenya" pageId="55">Kenya</collectingCountry>
, 
<typeStatus id="54A3881CFFB1FFC9FCA93ACA7C7AFC46" box="[771,819,921,945]" pageId="55">type</typeStatus>
species of the genus), 
<taxonomicName id="4C184D3DFFB1FFC9FBEE3ACA7C51FC26" authority="Chandler" authorityName="Chandler" authorityYear="1994" class="Insecta" family="Platypezidae" genus="Lindneromyia" kingdom="Animalia" order="Diptera" pageId="55" phylum="Arthropoda" rank="species" species="fonsecai">
<emphasis id="B96CEAACFFB1FFC9FBEE3ACA7BF5FC47" box="[1092,1212,920,944]" italics="true" pageId="55">L. fonsecai</emphasis>
Chandler
</taxonomicName>
(from 
<collectingCountry id="F30F762EFFB1FFC9FC2B3AEA7CA6FC26" box="[897,1007,953,977]" name="Australia" pageId="55">Australia</collectingCountry>
), and 
<taxonomicName id="4C184D3DFFB1FFC9FBE63AE97C4CFC05" authority="Bowden" authorityName="Bowden" authorityYear="1973" class="Insecta" family="Platypezidae" genus="Lindneromyia" kingdom="Animalia" order="Diptera" pageId="55" phylum="Arthropoda" rank="species" species="kesseli">
<emphasis id="B96CEAACFFB1FFC9FBE63AE97BF5FC26" box="[1100,1212,953,977]" italics="true" pageId="55">L. kesseli</emphasis>
Bowden
</taxonomicName>
(from 
<collectingCountry id="F30F762EFFB1FFC9FCFB3A897CA9FC05" box="[849,992,986,1010]" name="South Africa" pageId="55">South Africa</collectingCountry>
). 
<taxonomicName id="4C184D3DFFB1FFC9FC5B3A897C41FBE4" authorityName="Grimaldi" authorityYear="2018" class="Insecta" family="Platypezidae" genus="Lindneromyia" kingdom="Animalia" order="Diptera" pageId="55" phylum="Arthropoda" rank="species" species="neomedialis">
<emphasis id="B96CEAACFFB1FFC9FC5B3A897C41FBE4" italics="true" pageId="55">Lindneromyia neomedialis</emphasis>
</taxonomicName>
significantly differs from these three species based on the single pair of dorsocentrals (vs. in two rows) and scutellars (vs. 2–3 pairs); lack of stiff setae on tergite 6 (which is widespread in 
<taxonomicName id="4C184D3DFFB1FFC9FCBD3D2C7CECFB60" box="[791,933,1151,1175]" class="Insecta" family="Platypezidae" kingdom="Animalia" order="Diptera" pageId="55" phylum="Arthropoda" rank="family">Platypezidae</taxonomicName>
); and the very distinctive, short dm-cu crossvein distant from the wing margin (vs. very close to the margin). 
<taxonomicName id="4C184D3DFFB1FFC9FBF83D917C01FB0C" authorityName="Chandler" authorityYear="1994" class="Insecta" family="Platypezidae" genus="Lindneromyia" kingdom="Animalia" order="Diptera" pageId="55" phylum="Arthropoda" rank="species" species="fonsecai">
<emphasis id="B96CEAACFFB1FFC9FBF83D917C01FB0C" italics="true" pageId="55">Lindneromyia fonsecai</emphasis>
</taxonomicName>
has a very abrupt bend in M 
<subScript id="179C34FBFFB1FFC9FB193DBD7BF5FB08" attach="left" box="[1203,1212,1262,1279]" fontSize="7" pageId="55">1</subScript>
(some individuals retain a spur of M 
<subScript id="179C34FBFFB1FFC9FBF73C5C7B2FFAD7" attach="right" box="[1117,1126,1295,1312]" fontSize="7" pageId="55">2</subScript>
). 
<taxonomicName id="4C184D3DFFB1FFC9FB2B3C577C3CFACA" class="Insecta" family="Platypezidae" genus="Lindneromyia" kingdom="Animalia" order="Diptera" pageId="55" phylum="Arthropoda" rank="species" species="africana">
<emphasis id="B96CEAACFFB1FFC9FB2B3C577C3CFACA" italics="true" pageId="55">Lindneromyia africana</emphasis>
</taxonomicName>
is reported to lack the small basal aristomeres (which would be very unusual, perhaps an observational error), and the basal flagellomere is setose (not just with microtrichia); dm-cu is extremely close to the wing margin (~ 0.3× crossvein length), and R 
<subScript id="179C34FBFFB1FFC9FBF53C857B21FA10" attach="left" box="[1119,1128,1494,1511]" fontSize="7" pageId="55">1</subScript>
is quite long (terminating at level of CuA 
<subScript id="179C34FBFFB1FFC9FBB93CA47B55F9FF" attach="left" box="[1043,1052,1527,1544]" fontSize="7" pageId="55">1</subScript>
apex). Despite these differences I am hesitant to accord a separate generic status to the fossil, especially since the external male terminalia agree so closely with 
<taxonomicName id="4C184D3DFFB1FFC9FD4F3F3C7C35F970" authorityName="Kessel" authorityYear="1965" box="[741,892,1647,1671]" class="Insecta" family="Platypezidae" genus="Lindneromyia" kingdom="Animalia" order="Diptera" pageId="55" phylum="Arthropoda" rank="genus">
<emphasis id="B96CEAACFFB1FFC9FD4F3F3C7C35F970" box="[741,892,1647,1671]" italics="true" pageId="55">Lindneromyia</emphasis>
</taxonomicName>
(e.g., 
<bibRefCitation id="EF894B4FFFB1FFC9FC6A3F3C7B22F97F" author="Chandler, P. J." box="[960,1131,1647,1672]" pageId="55" pagination="351 - 434" refId="ref38631" refString="Chandler, P. J. 1994. The Oriental and Australasian species of Platypezidae (Diptera). Invertebrate Systematics 8: 351 - 434." type="journal article" year="1994">Chandler, 1994</bibRefCitation>
; 
<bibRefCitation id="EF894B4FFFB1FFC9FBDC3F3C7C6BF95E" author="Chandler, P. J." pageId="55" pagination="1 - 276" refId="ref38658" refString="Chandler, P. J. 2001. The flat-footed flies (Diptera: Opetiidae and Platypezidae) of Europe. Fauna Entomologica Scandinavica 36: 1 - 276. Leiden: Brill." type="journal article" year="2001">Chandler, 2001</bibRefCitation>
). Also, many of the distinctive features of 
<taxonomicName id="4C184D3DFFB1FFC9FCAE3FE17CD4F93D" authorityName="Grimaldi" authorityYear="2018" box="[772,925,1714,1738]" class="Insecta" family="Platypezidae" genus="Lindneromyia" kingdom="Animalia" order="Diptera" pageId="55" phylum="Arthropoda" rank="species" species="neomedialis">
<emphasis id="B96CEAACFFB1FFC9FCAE3FE17CD4F93D" box="[772,925,1714,1738]" italics="true" pageId="55">L. neomedialis</emphasis>
</taxonomicName>
are autapomorphic losses.
</paragraph>
<caption id="DF676636FFBEFFC6FFC73C8D7B04F9E5" ID-DOI="http://doi.org/10.5281/zenodo.4613096" ID-Zenodo-Dep="4613096" httpUri="https://zenodo.org/record/4613096/files/figure.png" pageId="56" startId="56.[109,151,1502,1523]" targetBox="[186,1116,231,1467]" targetPageId="56">
<paragraph id="8BA736BEFFBEFFC6FFC73C8D7B04F9E5" blockId="56.[109,1213,1500,1554]" pageId="56">
FIG. 35. 
<taxonomicName id="4C184D3DFFBEFFC6FF6E3C8F7E80FA03" authority="Grimaldi, 2018" authorityName="Grimaldi" authorityYear="2018" box="[196,457,1500,1524]" class="Insecta" family="Platypezidae" genus="Lindneromyia" kingdom="Animalia" order="Diptera" pageId="56" phylum="Arthropoda" rank="species" species="dominicana" status="sp. nov.">
<emphasis id="B96CEAACFFBEFFC6FF6E3C8F7E80FA03" box="[196,457,1500,1524]" italics="true" pageId="56">Lindneromyia dominicana</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A25F57D7FFBEFFC6FE7F3C8E7D43FA03" box="[469,522,1501,1524]" pageId="56" rank="species">n. sp.</taxonomicNameLabel>
(
<taxonomicName id="4C184D3DFFBEFFC6FDBC3C8F7DD2FA03" box="[534,667,1500,1524]" class="Insecta" family="Platypezidae" kingdom="Animalia" order="Diptera" pageId="56" phylum="Arthropoda" rank="family">Platypezidae</taxonomicName>
), male holotype in Dominican amber, AMNH 14-736. 
<emphasis id="B96CEAACFFBEFFC6FFC73CAA7FCFF9E6" bold="true" box="[109,134,1529,1553]" pageId="56">A.</emphasis>
Wing. 
<emphasis id="B96CEAACFFBEFFC6FF783CA97FA0F9E5" bold="true" box="[210,233,1530,1554]" pageId="56">B.</emphasis>
Apex of abdomen, left lateral. 
<emphasis id="B96CEAACFFBEFFC6FD8E3CAA7D75F9E6" bold="true" box="[548,572,1529,1553]" pageId="56">C.</emphasis>
Protarsomeres 1–4. 
<emphasis id="B96CEAACFFBEFFC6FCA43CA97C69F9E5" bold="true" box="[782,800,1530,1554]" pageId="56">D</emphasis>
. Metatarsomeres 1–3, lateral.
</paragraph>
</caption>
<paragraph id="8BA736BEFFBFFFC7FF2639B17FB7FDF5" blockId="57.[108,638,226,514]" pageId="57">
<taxonomicName id="4C184D3DFFBFFFC7FF2639B17EE1FF0D" authorityName="Grimaldi" authorityYear="2018" box="[140,424,226,250]" class="Insecta" family="Platypezidae" genus="Lindneromyia" kingdom="Animalia" order="Diptera" pageId="57" phylum="Arthropoda" rank="species" species="neomedialis">
<emphasis id="B96CEAACFFBFFFC7FF2639B17EE1FF0D" box="[140,424,226,250]" italics="true" pageId="57">Lindneromyia neomedialis</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="4C184D3DFFBFFFC7FD9639B17FFAFEEC" authorityName="Kessel" authorityYear="1952" class="Insecta" family="Platypezidae" genus="Metaclythia" kingdom="Animalia" order="Diptera" pageId="57" phylum="Arthropoda" rank="genus">
<emphasis id="B96CEAACFFBFFFC7FD9639B17FFAFEEC" italics="true" pageId="57">Metaclythia</emphasis>
</taxonomicName>
(monotypic: 
<emphasis id="B96CEAACFFBFFFC7FEE338507EDEFEED" box="[329,407,259,282]" italics="true" pageId="57">currani</emphasis>
Kessel, known from 
<collectingRegion id="49DCF85CFFBFFFC7FFC638777F8DFECB" box="[108,196,292,316]" country="Canada" name="Ontario" pageId="57">Ontario</collectingRegion>
, 
<collectingCountry id="F30F762EFFBFFFC7FF6438777E6AFECB" box="[206,291,292,316]" name="Canada" pageId="57">Canada</collectingCountry>
, and 
<collectingRegion id="49DCF85CFFBFFFC7FEF438777E9AFECB" box="[350,467,292,316]" country="United States of America" name="Wisconsin" pageId="57">Wisconsin</collectingRegion>
, 
<collectingCountry id="F30F762EFFBFFFC7FE7738777D39FECB" box="[477,624,292,316]" name="United States of America" pageId="57">United States</collectingCountry>
), which also has a simple M vein, based on the following: Cell dm short (vs. long), gena bare (vs. setose); single pair of dorsocentrals (vs. in graded row); scutellars in single pair (vs. two); 2 (vs. 4) notopleurals; tergite 6 lacking spinelike setae (vs. with 2 pairs).
</paragraph>
</subSubSection>
</treatment>
</document>