Three new genera and eighteen new species of miniature polydesmid millipedes from the northwestern United States (Diplopoda, Polydesmida, Polydesmidae)
Author
Shear, William A.
Professor Emeritus, Department of Biology, Hampden-Sydney College, Hampden-Sydney VA 23943; present address 1950 Price Drive, Farmville VA 23901.
Author
Marek, Paul E.
0000-0002-7048-2514
Associate Professor, Department of Entomology, Virginia Polytechnic Institute and State University, Blacksburg, VA 24061. pmarek @ vt. edu; https: // orcid. org / 0000 - 0002 - 7048 - 2514
pmarek@vt.edu
text
Zootaxa
2021
2021-05-24
4975
1
81
126
journal article
6194
10.11646/zootaxa.4975.1.3
529d48db-4299-4e2d-9aaf-6c4178c08b18
1175-5326
4804909
DB7C9028-3EDF-454F-88D0-336624AD1DC4
Retrorsioides bammerti
Shear & Marek
,
new species
Figs 76–78
,
130
Types.
Male
holotype
and male
paratypes
from
Bob Bammert Grove
,
Capitol State Forest
,
287 ft
asl
, 46.8960°, - 123.0963°,
Thurston Co.
,
Washington
, collected
14 November 2004
by
W. Leonard. All
types deposited in
CAS
.
Diagnosis.
The sigmoid terminal zone of the gonopod (
tz,
Fig. 130
) and the basal position of the pulvillus (
p,
Fig. 130
) set this species apart from all others in the genus.
Etymology.
The species name recognizes Robert (Bob) Bammert (1926–2019), noted
Washington
conservationist for whom the
type
locality, a grove of old-growth trees, is named.
Description.
Male
holotype
. Length about 4.0 mm, greatest width
0.50 mm
. Head densely setose, cuticle alveolate (
Fig. 76
). Collum (
Fig. 76
) with anterior marginal row of 14 setae. Anterior metazonites with three rows of setae (
Fig. 76
), transitioning to four rows (
Fig. 77
) about ring 5, posteriormost segments with five rows (
Fig. 78
); setal tubercles becoming almost obsolete on midbody to posterior rings. Alveolate cuticle absent from metazonites posterior to collum. Epiproct (
Fig. 78
) swollen, short, bent ventrally, without alveolate cuticle. Anterior legs crassate, tarsi with sphaerotrichomes. Gonopod with oblong, inflated prefemorite (
pf,
Fig. 130
) similar to previous species. Acropodite short, stout. Pulvillus (
p,
Fig. 130
) in basal third of acropodite, pulvillar process (
pp,
Fig. 130
) long, acute, directed distally. Anteriorly directed process (
adp,
Fig. 130
) rounded, tapering slightly, without teeth. Terminal zone (
tz,
Fig. 130
) not bifurcate, sigmoidally curved. Females unknown.
Distribution.
Northwestern
Washington
in Thurston, Pacific and Lewis Counties.
Records:
WASHINGTON
:
Lewis Co.
: Stillman Basin,
1.8 mi
on Weyerhauser 4200 from road W4000, 2116 ft asl, 48.4956°, -123.2134°,
4 December 2004
, W. Leonard, C. Richart, mm.
Pacific Co.
:
1.1 mi
south of SR6 on Trap Creek Road, B-line, 46.5432°, -123/6152°, W. Leonard, C. Richart, m.
Note:
At the Stillman Basin locality, this species is syntopic with
Retrorsia simplicissima
.