A revision of the genus Rhinotorus Förster, 1869 (Hymenoptera, Ichneumonidae, Ctenopelmatinae), with descriptions of three new species and an illustrated identification key
Author
Reshchikov, Alexey V.
text
European Journal of Taxonomy
2016
2016-10-05
235
235
1
40
journal article
21870
10.5852/ejt.2016.235
6c2c2795-d283-4de4-a94e-42c64c88187c
2118-9773
3847734
1A763AF4-A12F-4CCF-94CB-3981D29D7E55
Genus
Rhinotorus
Förster, 1869
Rhinotorus
Förster, 1869: 211
.
Spudaea
Förster, 1869: 211
Preocc.
Spudaeus
Thomson, 1883: 932
Preocc.
Prospudaea
Hincks, 1944: 35
.
Type
species:
Spudea longicornis
Schmiedeknecht, 1914
; designated by
Perkins (1962)
.
Diagnosis
(updated from Townes 1970)
Small to moderate sized species, fore wing length
4.5–9 mm
. Ground colour mainly black. Antenna slender to stout, with 28–38 flagellomeres. Scape 1.25–1.6 times as long as broad. Face usually convex and densely punctate. Clypeus small, convex, around 0.3–0.4 times as long as broad, distinctly separated from face by a deep impression, medially projecting, shining and sparsely punctate, its apical margin rounded, laterally impressed and sharp. Malar space 0.5–0.7 times basal mandible width. Occipital carina complete. Mandible with teeth usually of equal length. Notaulus distinct. Mesopleuron usually granulate, striated, or densely and coarsely punctate. Metapleuron weakly shining, rugose, with rather dense punctation in upper part. Propodeum strongly rugose. Area superomedia usually fused with area basalis. Costula absent. Area apicalis large, trapezoidal, often with longitudinal carina and sometimes with striation. Spurs of middle and hind tibiae of unequal length. Tarsal claws not pectinate. Fore wing with pterostigma elongate. 2m-cu straight, with a single bulla. Cu-a vertical, slightly postfurcal. Hind wing with 1/Cu & cu-a intercepted by 2/Cu in lower 0.5. Metasoma coarsely punctate and reticulate rugose. First metasomal tergite usually short and projecting dorsally, its dorsal longitudinal carinae reaching middle of tergite, with a strong impression between carinae in basal part. Subapical impression strong. Second metasomal tergite with transverse impression starting immediately after middle of tergite. Third tergite with weak transverse impression. Ovipositor sheaths weakly clavate. Ovipositor stout at base with triangular shape before shallow subapical notch, its lower valvae slightly swollen before thin tip and tip of upper valve rather elongate after notch. Parameres broad basally and weakly elongate apically, extending beyond aedeagus. Volsella moderately large and notched apically. Aedeagus apically downcurved.
Tribal diagnosis
The genus belongs to the tribe
Mesoleiini
, whose representatives can be distinguished from other ctenopelmatines by combination of the following characters: epicnemial carina reaching fore edge of
Fig. 1. A–D
.
Saotis brachycerus
(Kasoparyan & Kopelke, 2009)
comb. nov.
A
. Face.
B
. Propodeum.
C
. Habitus paratype, Ƌ
S. brachycerus
(Kasoparyan & Kopelke, 2009)
.
D
. Habitus holotype, ♀
S.brachycerus
(Kasoparyan & Kopelke, 2009)
.
E–F
.
Saotis granulator
Kasparyan & Kopelke, 2010
.
E
. Habitus Ƌ
S. granulator
Kasparyan & Kopelke, 2010
.
F
. Habitus ♀
S. granulator
Kasparyan & Kopelke, 2010
.
Fig. 2.
Mesoleius roepkei
Teunissen, 1945
, holotype, ♀.
A
. Face.
B
. First metasomal tergite with shallow subapical transverse impressions (arrow).
C
. Habitus.
mesopleuron, notch between basal part of propodeum (at its middle) and hind edge of postscutellum V-shaped, glymma present, claws mostly without pecten.
Species in the genus
Rhinotorus
can be recognized by combination of the following characters: first and second metasomal tergites with subapical transverse impressions. Metasoma strongly punctate and reticulate rugose. Median dorsal carinae of first metasomal tergite distinct. 2m-cu with a single bulla. Areolet absent. Glymma present. Hind wing with 1/Cu & cu-a intercepted by 2/Cu in lower 0.5.
Some representatives of other
Mesoleiini
genera also have subapical transverse impressions of metasomal tergites [e.g.,
Mesoleius roepkei
Teunissen, 1945
and
Saotis brachycerus
(Kasoparyan & Kopelke, 2009)
comb. nov.
(
Figs 1
,
2
C–D)], but these have a broad and short ovipositor sheath, or a compressed metasoma as in
Saotis
(
Fig. 1
).
Saotis brachycerus
(Kasoparyan & Kopelke 2009)
comb. nov.
originally was described in the genus
Rhinotorus
with note “
Saotis
” in the brakes (Kasoparyan & Kopelke 2009). However it should be considered as
Saotis
since it has the ovipositor sheaths shortly elliptic but not elongate cone- or club-shaped as in
Rhinotorus
.