A review of the genus Gynaecoserica Brenske, 1896 (Coleoptera, Scarabaeidae, Sericini) Author Ahrens, Dirk Author Fabrizi, Silvia text Journal of Natural History 2009 2009-06-16 43 25 - 26 1505 1584 http://dx.doi.org/10.1080/00222930902968809 journal article 10.1080/00222930902968809 1464-5262 5216589 Gynaecoserica variipennis wuermlii ( Frey, 1975 ) ( Figure 9B ) Lasioserica wuermlii Frey, 1975: 231 . Gynaecoserica variipennis wuermlii : Ahrens 1996: 37 ; 1999: 184 ; 2004: 107 . Remarks The species was revised by Ahrens (1996) . This subspecies of G. variipennis is distributed in Bhutan ( Figure 9B ).