A new genus and species of hawker dragonfly of uncertain affinities from the Middle Jurassic of China (Odonata: Aeshnoptera) Author Li, Yongjun Author Nel, André Author Ren, Dong Author Pang, Hong text Zootaxa 2011 2927 57 62 journal article 46580 10.5281/zenodo.201571 173b468a-5276-41c6-a05c-76ec6e7a8bc7 1175-5326 201571 Sinocymatophlebiella hasticercus sp. nov. ( Figs 1 –4) Etymology. Named after Latin prefix ‘hasti-’ and cerci, for our specimen has very strange hastate cerci. FIGURE 1. (a) Photograph of Sinocymatophlebiella hasticercus gen. et sp. nov. CNU-ODO-NN2010004-1, part. Scale bar represents 10 mm. (b) Photograph of Sinocymatophlebiella hasticercus gen. et sp. nov. under alcohol. CNU-ODO- NN2010004-2, counterpart. Scale bar represents 10 mm. FIGURE 2. Line drawing of Sinocymatophlebiella hasticercus gen. et sp. nov. Scale in mm. Type locality and horizon. Jiulongshan Formation, Middle Jurassic; near Daohugou Village, Wuhua Township, Ningcheng County, Inner Mongolia , China Diagnosis. As for the genus. Description. Forewing hyaline, preserved wing length 45.1 mm , 10.3 mm wide; distance between base and arculus 5.3 mm , between arculus and nodus circa 17.8 mm ; distance from nodus to wing apex circa 20.0 mm; distance between Ax1 and Ax2 5.2 mm , between Ax1 and wing base 3.6 mm ; five preserved secondary antenodal cross-veins, three of them being between Ax1 and Ax2; Ax2 lies at level of distal third of discoidal triangle; arculus straight; pterostigma and postnodal area not preserved; median space free; submedian space only traversed by CuP-crossing; PsA weakly zigzagged; hypertriangle free; discoidal triangle elongated, divided into three cells; MAb straight, 4.3 mm long; a well-defined three-celled subdiscoidal triangle; base of IR2 circa 3.7 mm basal of nodus; that of RP 3/4 circa 6.0 mm basal of nodus; RP 3/4 and MA parallel and distinctly undulated, with one row of cells between them basally and five rows of cells distally at their preserved length; MA and MP are more or less parallel, postdiscoidal area weakly widened at level of nodus; no Mspl; MP and CuA basally parallel with one row of rectangular cells between them, distally divergent; CuA with eight posterior branches, cubito-anal area with six rows of cells between CuA and posterior wing margin; two rows of cells in anal area; a very short subdiscoidal veinlet, 0.2 mm long. FIGURE 3. Photograph of abdomen segment II, scale in mm. FIGURE 4. Photograph of cerci, scale in mm. Hind wing hyaline, pterostigma dark brown; wing 42.6 mm long, 14.1 mm wide; distance between base and arculus 4.8 mm , between arculus and nodus 14.0 mm; distance from nodus to mid of pterostigma 16.6 mm ; distance between Ax1 and Ax2 5.5 mm , between Ax1 and wing base 3.7 mm ; six secondary antenodal cross-veins, two of them being between Ax1 and Ax2; Ax2 lies opposite distal angle of discoidal triangle; arculus straight; pterostigma elongated, 3.8 mm long and 0.7 mm wide, covering two cells, not basally recessed; pterostigmal brace slightly oblique and aligned with basal side of pterostigma; eight postnodal cross-veins not aligned with postsubnodal cross-veins; median space free of cross-veins; submedian space only traversed by CuP-crossing; PsA straight; hypertriangle free; discoidal triangle elongated, divided into two cells; MAb straight, about 4.5 mm long; a welldefined two-celled subdiscoidal triangle; base of IR2 3.5 mm basal of nodus; that of RP 3/4 5.1 mm basal of nodus; only one cross-vein between RP and IR2 basal of first ‘O’, and this cross-vein is Bqs; two oblique veins ‘O’, about 1.7 mm and 6.1 mm distal of subnodus, second ‘O’ much more oblique than first one; pseudo-IR1 short, 1.7 mm distal of pterostigma; area between RP 1 and RP 2 rather narrow with one rows of cells between them in basal part; base of RP 2 well aligned with subnodus, RP 2 smoothly undulate at its mid part; IR2 weakly undulate, area between it and RP 2 widened at their undulate parts; IR2 and RP 2 basally parallel, with only one row of cells and three to six rows distally; a not very well-developed and zigzagged Rspl with one row of cells between it and IR2; no strongly convex oblique and undulating secondary vein anastomosing between IR2 and RP 3/4 directly basal of origin of Rspl; RP 3/4 and MA parallel and strongly undulate, with one row of cells between them basally and four rows distally; MA and MP more or less parallel, postdiscoidal area weakly widened at level of nodus; no Mspl; area between MP and CuA with one row of cells basally and distally divergent; CuAa with seven well-defined and parallel posterior branches; CuAb directed towards posterior wing margin, anal loops nearly absent, posteriorly completely opened, gaff (basal part of CuA) short, 0.6 mm long; two posterior branches of AA between distal side of anal triangle and CuAb; a distinct anal angle and a three-celled anal triangle (male specimen); a long and broad, strongly sclerotized membranule. Body: total body length (from head up to end of abdomen, including anal appendages) 83.6 mm . Head 5.8 mm long, 6.2 mm wide, with compound eyes large, medially contiguous; thorax 6.6 mm wide with weak traces of coloration; fore legs partly preserved, tibia about 6.3 mm long, tarsi not visible, femur about 5.7 mm long; abdomen 57.5 mm long, the most wide area 5.2 mm ; first abdominal segment 2.2 mm long, second 12.1 mm long, third 7.4 mm long; second segment distinctly constricted with a width of 2.0 mm, no auricles visible on second segment ( Fig. 3 ); no pair of symmetrical expanded and rounded lobes on abdominal segments III or IV; cerci clearly visible, narrow, 6.1 mm long and about 1.0 mm wide, with strong apical and outer spines and a pronounced dark line suggesting the presence of a longitudinal crest (Fig. 4); inferior appendage (epiproct) very small (about 0.5 mm long), hardly visible.