Prodromus of a fern flora of Bolivia. XLII. Update I.
Author
Kessler, Michael
Systematic and Evolutionary Botany, University of Zurich, Zollikerstrasse 107, CH- 8008 Zurich, Switzerland.
Author
Smith, Alan R.
University Herbarium, 1001 Valley Life Sciences Bldg. # 2465, University of California, Berkeley, CA 94720 - 2465.
Author
Øllgaard, Benjamin
Department of Biology - Ecoinformatics and Biodiversity, Ny Munkegade 116, building 1137, 8000 Aarhus C, Denmark.
Author
Matos, Fernando B.
Centro de Referência em Informação Ambiental (CRIA), Rua João Carlos do Amaral 500, 13070 - 111, Campinas, SP, Brasil.
Author
Moran, Robbin C.
New York Botanical Garden, 2900 Southern Blvd., Bronx, New York, 10458 - 5126, USA.
text
Phytotaxa
2023
2023-12-20
630
3
183
210
https://phytotaxa.mapress.com/pt/article/download/phytotaxa.630.3.2/51376
journal article
10.11646/phytotaxa.630.3.2
1179-3163
10409920
Parrisia
Shalisko & Sundue, Phytotaxa
394(3): 187. 2019
.
The grammitid genus
Parrisia
was named recently in a paper by
Shalisko
et al
. (2019)
to accommodate two species previously treated in
Enterosora
by
Bishop & Smith (1992)
,
Smith & Bishop (1995)
, and Smith
et al
. (2018).
Parrisia parietina
is widespread, from southern
Mexico
to
Bolivia
; a second species,
P. gilpinae
(Baker) Shalisko & Sundue
, is in
Tanzania
and
Madagascar
, and was treated under the name
E. sprucei
(Hook.) Parris (
Parris 2005
)
, which we consider a synonym of
P. parietina
.
This unusual distribution pattern of
Parrisia
is also known in several other grammitid genera, e.g.,
Ceradenia
,
Enterosora
,
Grammitis
s.s
.
,
Leucotrichum
,
Melpomene
,
Moranopteris
, and
Stenogrammitis
.
Previously, species in
Parrisia
have generally been placed either in a very broadly defined genus
Grammitis
, a definition that included all or nearly all grammitid ferns (700+ spp.), or in
Enterosora
(
Bishop & Smith 1992
; Smith
et al
. 2018).
Parrisia
agrees with
Ceradenia
,
Enterosora
,
Grammitis
s.s
.
, and several other grammitid genera in lacking clearly defined hydathodes adaxially. From
Enterosora
,
Parrisia
is distinguished by the radial rhizomes (vs. dorsiventral), the sori borne at the middle of veins (vs. distally), and the presence of setae around the sori (
Shalisko
et al
. 2019
). From other simple-bladed New World grammitids, e.g.,
Cochlidium
,
Parrisia
differs in lacking hydathodes, the presence of dark-colored setae on the blades, and from some species by the superficial sori (vs. immersed coenosori). Another simple-bladed genus in the Neotropics is
Lomaphlebia
, endemic to the Greater Antilles; that genus has a row of marginal areoles on the blades.
Grammitis
s.s
.
, also simple-bladed, differs in having black sclerotized blade margins and general lack of setae abaxially on the blades.
Phylogenetic studies suggest that
Parrisia
may be related to the endemic Hawaiian genus
Adenophorus
, with species having distinctive reddish glandular paraphyses in the sori and on the lamina abaxially, to Antillean
Lomaphlebia
(2 spp.), and to the amphioceanic genera
Grammitis
s.s
.
and
Cochlidium
(all but one species neotropical). Affinities with
Enterosora
(see above; now including
Zygophlebia
) are more remote (
Bauret
et al
. 2017
;
Shalisko
et al
. 2019
). The current name for the Bolivian species is: