New or rare Madagascar tiger beetles- 23. Physodeutera (Toxoma) lokobensis sp nov., a new species close to Ph. (T.) conturbata Moravec, Ph. (T.) sulcoprothoracica (W. Horn) and Ph. (T.) dubia (Mařan), with revised type designation of the latter (Coleoptera: Cicindelidae) Author Moravec, Jiří Author Trýzna, Miloš text Zootaxa 2021 2021-10-29 5060 2 151 182 journal article 3777 10.11646/zootaxa.5060.2.1 ed98a754-c3ac-49ff-9973-b8d9839993ff 1175-5326 5627213 F6BFFABC-7F88-473C-9517-D031485140DD Key to species of Physodeutera , subgenus Toxoma (individual females may be barely identifiable in some species) 1 Apex of antennomere 4 dilated in male. Elytra entirely immaculate in both sexes. Body medium-sized, bright metallic blue with green, greenish-bronze and violet reflections. Aedeagus long with elongate basal half, apical portion ventrally arcuate-emarginated, forming ventrad-directed, rounded apex............................................................... 2 - Antennomere 4 normally shaped. Body medium-sized to large. Elytra with at least humeral macula in male.Aedeagus variously shaped.............................................................................................. 3 2 Antennomere 4 (in male) simply clavate-dilated. Lateral margins of pronotal disc almost parallel, slightly emarginated in middle. Pronotum slightly more elongate with rather rounded, less parallel lateral margins of disc. Elytra bright green-blue. Aedeagus moderately enlarged in middle.............................. Ph . ( T .) lobicornis lobicornis Moravec, 2000 - Antennomere 4 (in male) more distinctly dilated, eccentric-spatulate. Elytra deep green-blue. Aedeagus more distinctly enlarged in middle.................................................. Ph . ( T .) lobicornis nosybensis Moravec, 2000 3 Elytra with conspicuous or well noticeable velvety black shadowy zone on elytral disc; punctures rather coarse and occasionally anastomosing on anterior elytral half, much finer or almost effaced on posterior elytral half...................... 4 - Elytra without noticeable velvety black zone on elytral disc (yet inconspicuous shadowy areas may diffuse on elytral disc or various elytral areas)................................................................................... 5 4 Body medium sized. Elytral disc with conspicuous, well-delimited velvety black shadowy zone in middle, remaining surface metallic black with feeble or bright, blue, green-blue, or bronze reflections, or with chatoyant, gold-bronze, or purple lustre on posterior declivity; whitish maculation consisting of humeral-lateral lunule (absent in female) and sublateral-median macula. Antennomere 4 in male ochre testaceous with black-brown base and apical third. Aedeagus short, attenuated into slightly ventrad-bent, rather wide, rounded apex...................................... Ph . ( T .) subtilevelutina ( W. Horn, 1934 ) - Body larger. Elytral disc with faint, yet notable velvety-black shadowy zone diffusing from central area towards outer elytral margins; remaining surface deep cyaneous-blue and green-blue with strong violaceous lustre laterally, sometimes with indistinct bronze lustre; only whitish humeral macula in male, which is in female only indicated as a small, dark, and almost invisible spot, or entirely absent. Antennomere 4 black with basal half light brownish or dark brown. Aedeagus with notably short apical portion towards very short, regularly rounded apex........................ Ph . ( T .) sulcoprothoracica (W. Horn, 1913 ) 5 Elytra with only white humeral macula in male, female elytra immaculate or with only inconspicuous, darkened humeral macula. Aedeagus rather stout (including its basal portion).................................................... 6 - Elytra also with white sublateral-median macula, punctate throughout, punctures coarser and anastomosing on anterior elytral half, much finer on posterior half. Aedeagus with narrowly elongate basal half..................................... 8 6 Antennomere 4 in male with two basal thirds ivory to ochre-testaceous, elytra bright green-blue, only lateral areas and apices violaceous........................................................................................... 7 - Antennomere 4 entirely black or black-brown in both sexes. Elytra rather coarsely punctate on their anterior half, posterior half with finer and notably spaced punctures yet not entirely effaced, deep cyaneous-violaceous with brighter greenish and indistinct bronze lustre within discal impression; elytral disc with only barely noticeable, feebly diffusing shade. Pronotal disc distinctly attenuated anteriad in male. Aedeagus apex short, regularly rounded.................. Ph . ( T .) conturbata Moravec, 2002 7 Elytra distinctly, almost equally punctate throughout, lacking any shadowy area on elytral disc. Pronotal disc less notably attenuated anteriad in male. Aedeagus apex regularly rounded yet narrower. Female unknown.... Ph . ( T .) dubia ( Mařan, 1942 ) - Elytral punctation finer, particularly so and spaced on posterior half of elytral disc, sometimes with indistinct shade in middle. Aedeagus apex wider, rounded but almost triangular-topped (ventrally moderately arcuate and dorsally obliquely sloped towards indistinct emargination)..................................................... Ph . ( T .) lokobensis sp. nov. 8(5-) Pronotum elongate. Elytra predominantly metallic green to green-blue, punctures coarser on anterior elytral half; whitish humeral macula combined with subhumeral-lateral lunule in male, while in female only subhumeral lunule present. Aedeagus with notably narrowly elongate basal half, dorsally enlarged apical half, while ventral margin straight; internal sac notably simple.................................................................... Ph . ( T .) sikorai (W. Horn, 1896) - Pronotum approximately as long as wide with notably wider disc. Elytra predominantly deep cyaneous, distinctly punctate throughout; whitish humeral macula in male simple, absent in female, yet obscurely indicated subhumeral appendage may occur in both sexes; lateromedian macula smaller. Aedeagus shorter, its apical half more enlarged and apex slightly directed ventrad; internal sac more armed, with distinct, longitudinal-sigmoid ventral piece with clavate apex......................................................................................... Ph . ( T .) breviformis (W. Horn, 1904)