Order Carnivora Author Wilson, Don E. Author Reeder, DeeAnn text 2005 The Johns Hopkins University Press Baltimore Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 1 532 628 book chapter 0-8018-8221-4 10.5281/zenodo.7316519 Lycalopex Burmeister 1854 Lycalopex Burmeister 1854 , Systematische Uebersicht der Thiere Brasiliens: 95-101 . Type Species: Canis magellanicus Gray 1847 Synonyms: Angusticeps Hilzheimer 1906 ; Eunothocyon J. A. Allen 1905 ; Lupulus Trouessart 1897 ; Microcyon Trouessart 1906 ; Nothocyon Wortman and Matthew 1899 ; Pseudalopex Burmeister 1856 ; Pseudolopex Philippi 1903 ; Pseudolycos Philippi 1903 ; Thous Gray 1869 ; Viverriceps Hilzheimer 1906 . Species and subspecies: 6 species with 11 subspecies: Species Lycalopex culpaeus ( Molina 1782 ) Subspecies Lycalopex culpaeus subsp. culpaeus Molina 1782 Subspecies Lycalopex culpaeus subsp. andinus Thomas 1914 Subspecies Lycalopex culpaeus subsp. lycoides Philippi 1896 Subspecies Lycalopex culpaeus subsp. magellanicus Gray 1837 Subspecies Lycalopex culpaeus subsp. reissii Hilzheimer 1906 Subspecies Lycalopex culpaeus subsp. smithersi Thomas 1914 Species Lycalopex fulvipes (Martin 1837) Species Lycalopex griseus ( Gray 1837 ) Species Lycalopex gymnocercus (G. Fischer 1814) Subspecies Lycalopex gymnocercus subsp. gymnocercus G. Fischer 1814 Subspecies Lycalopex gymnocercus subsp. antiquus Ameghino 1889 Subspecies Lycalopex gymnocercus subsp. domeykoanus Philippi 1901 Subspecies Lycalopex gymnocercus subsp. gracilis Burmeister 1861 Subspecies Lycalopex gymnocercus subsp. maullinicus Philippi 1903 Species Lycalopex sechurae Thomas 1900 Species Lycalopex vetulus (Lund 1842) Discussion: Revised by Zunino et al. (1995) . Also see comments under Dusicyon . Although combining taxa included here with Dusicyon would not be in conflict with Berta (1987 , 1988 ), her analyses suggested that other genera, now extinct, are more closely related to Dusicyon . Berta (1987 , 1988 ) presented derived features that would support a single origin for those taxa recognized here in Lycalopex (= Pseudalopex ), which would also agree with Cabrera (1957) and Stains (1975) . A detailed comparative morphological study by Langguth (1969) caused him to conclude (1975) that Pseudalopex (= Lycalopex ) merited generic rank. Synonyms allocated according to Zunino et al. (1995) . Tedford et al. (1995) considered Lycalopex as recognized here as paraphyletic and proposed a different arrangement of South American canids according to the following monophyletic groups: (1) L. vetulus + Chrysocyon + Cerdocyon + Nyctereutes + Speothos + Atelocynus ; (2) L. culpaeus + Dusicyon ; (3) L. griseus + L. gymnocercus + L. sechurae .