Revision and cladistic analysis of the Southeast Asian leaf-dwelling spider genus Calapnita Simon (Araneae, Pholcidae)
Author
Bernhard A. Huber
text
Zootaxa
2017
4219
1
1
63
journal article
37319
10.5281/zenodo.273086
0d2332a5-1968-49d7-a570-f31d386eccb3
1175-5326
273086
0FA0F51A-3868-4F13-A93D-E34CA5A689F8
Calapnita bugis
sp. nov.
Figs 180–187
“
Calapnita vermiformis
” (misidentification; only specimens from
Sulawesi
):
Deeleman-Reinhold 1986a
: 45
; 1986b: 212.
Huber 2011
: 48
, figs 141, 147–152.
Diagnosis
. Distinguished from most species of
vermiformis
group (except
C. dayak
) by wide obtuse tip of ventral flap of procursus (
Figs 180–181
); from most (except
C. saluang
) also by thin proximal part of bipartite cheliceral apophysis (
Fig. 184
) and by strong and regular fringes at tip of embolus; from most (except
C. bohol
) also by oval pore plates in transversal orientation (
Fig. 187
); from several species also by narrow and distinct epigynal ‘knob’ (
Fig. 186
).
FIGURES 172–187
.
Calapnita lawangan
sp. nov.
(172–179; RMNH, ARA 17815–16) and
C. bugis
sp. nov.
(180–187; RMNH, ARA 17420).
172–173, 180–181
. Left procursi, prolateral and retrolateral views (arrow points at distinctive undulating ridge).
174, 182
. Left palpal trochanters, retrolateral views.
175, 183
. Left palpal femora, prolateral views.
176–177, 184–185
. Right male chelicerae, frontal and lateral views.
178, 186
. Epigyna, ventral views.
179, 187
. Cleared female genitalia, dorsal views. Scale lines: 0.3 mm.
Etymology
. Named for the Buginese people, an ethnic group in southwestern
Sulawesi
; noun in apposition.
Material examined.
Holotype
. INDONESIA-SULAWESI:
♂
,
RMNH
(
ARA
17817),
South Sulawesi
,
55 km
from
Udjung Pandang
,
23 km
from
Camba
[~
5.05°S
,
119.74°E
], under leaves in karst forest, 9 &
13–14.viii.1980
(
C.L. & P.R. Deeleman
).
Other material
. INDONESIA-SULAWESI:
3♂
3♀
,
RMNH
(
ARA
17420), same data as holotype
;
1♂
1♀
,
RMNH
(
ARA
17426), same data but only
9.viii.1980
.
2♂
3♀
,
RMNH
(
ARA
17423),
North Sulawesi
,
Dumoga
, near
Doloduo
[~
0.517°N
,
123.917°E
], primary forest, from leaves,
27–30.viii.1982
(collector not given).
Assigned tentatively
. INDONESIA-SULAWESI:
2♂
2♀
,
RMNH
(
ARA
17425),
Central Sulawesi
,
Palu
,
Marena
,
Lore Lindu Reserve
[~
1.3°N
, 120, 083°E], “dark little stream valley”, from leaves,
24.vii.1982
(collector not given).
Description. Male
(
holotype
)
MEASUREMENTS. Total body length 5.2, carapace width 0.65. Leg 1 missing, tibia 2: 5.5, tibia 3: 3.4, tibia 4: 6.0. Distance PME-PME 200 µm, diameter PME 80 µm, distance PME-ALE ~30 µm; no trace of AME.
COLOR. Entire animal very pale gray to whitish, legs with brown patellae and tibia-metatarsus joints.
BODY. Habitus as in close relatives (cf.
Figs 120, 122
); ocular area barely elevated, each triad on very low hump; carapace without median furrow; clypeus unmodified; sternum barely wider than long (0.48/0.46), unmodified.
CHELICERAE. As in
Figs 184–185
, apophyses near lamellae clearly bipartite, proximal parts slender (similar to
C. saluang
), with pair of barely visible lateral processes; without modified hairs; without stridulatory ridges.
PALPS. In general similar to
C. vermiformis
(cf. figs 139 and
140 in
Huber 2011
); trochanter apophysis slender, as in
Fig. 182
; femur as in
Fig. 183
, distal process at 44% of femur length; tibia length/diameter 0.51/0.25; procursus as in
Figs 180–181
, with simple ventro-distal flat sclerite; bulb length 0.40; embolus length 0.60; embolus tip with strong and regular semi-transparent fringes.
LEGS. Without spines and curved hairs; few vertical hairs.
Male
(variation). Tibia
1 in
three other males: 7.7, 7.7, 7.8. The specimens from Lore Lindu Reserve are assigned tentatively because the tip of the procursus differs slightly (ventro-distal flat sclerite directed slightly more towards ventral).
Female
. In general similar to male; eye triads slightly closer together (distance PME-PME 180 µm). Tibia
1 in
3 females
: 6.3, 6.5, 6.7. Epigynum as in
Fig. 186
, weakly sclerotized triangular plate with small and slender anterior ‘knob’; internal genitalia as in
Fig. 187
, much longer than wide, with small elongated pore plates in transversal orientation far from posterior margin.
Distribution.
Apparently widespread in
Sulawesi
(
Fig. 283
).