Revision and cladistic analysis of the Southeast Asian leaf-dwelling spider genus Calapnita Simon (Araneae, Pholcidae)
Author
Bernhard A. Huber
text
Zootaxa
2017
4219
1
1
63
journal article
37319
10.5281/zenodo.273086
0d2332a5-1968-49d7-a570-f31d386eccb3
1175-5326
273086
0FA0F51A-3868-4F13-A93D-E34CA5A689F8
Calapnita dayak
sp. nov.
Figs 164–171
“
Calapnita vermiformis
” (misidentification; only specimens listed below):
Deeleman-Reinhold 1986b
: 212
.
Huber 2011
: 48
.
Diagnosis
. Distinguished from most species of
vermiformis
group (except
C. bugis
) by wide obtuse tip of ventral flap of procursus (
Fig. 164
); from most (except
C. bario
) also by long and slender palpal tibia (tibia length/width>2.5;
Fig. 286
); from several species also by very small proximal part of bipartite cheliceral apophyses (
Fig. 168
), by narrow and distinct epigynal ‘knob’ (
Fig. 170
), and by oval pore plates converging anteriorly and very close together (
Fig. 171
).
Etymology
.
Dayak
is a loose term for over 200 riverine and hill-dwelling ethnic subgroups located principally in the interior of Borneo; noun in apposition.
Material examined.
Holotype
. INDONESIA-BORNEO:
♂
,
RMNH
(
ARA
17814),
Central Kalimantan
,
Tumbang Tahai
(
2.033°S
,
113.583°E
), primary marshy forest, from leaves, 3/
13.ix.1985
(
S. Djojosudharmo
).
Other material
. INDONESIA-BORNEO:
1♂
1♀
,
RMNH
(
ARA
17424), same data as holotype
.
1♂
1♀
,
RMNH
(
ARA
17421),
Central Kalimantan
,
Kaharian
(
2.033°S
,
113.667°E
), swampy primary forest, in foliage,
2– 16.ix.1985
(
S. Djojosudharmo
)
.
1♂
1♀
,
RMNH
(
ARA
17427),
Central Kalimantan
,
40 km
NW
Palangkaraya
[~
1.96°S
,
113.75°E
?], secondary forest, from leaves,
ix.1985
(
S. Djojosudharmo
).
Assigned tentatively
. INDONESIA-BORNEO:
1♂
1♀
,
ZFMK
(Ar 5330),
East Kalimantan
,
Balikpapan
[~
1.25°S
,
116.833°E
], primary forest,
20.vii.1982
(Murphy, 11872 part).
Description. Male
(
holotype
)
MEASUREMENTS. Total body length 3.5, carapace width 0.5. Leg 1: 25.4 (6.5 + 0.3 + 5.5 + 10.9 + 2.2), tibia 2: 4.2, tibia 3: 2.6, tibia 4: 4.7; tibia 1 L/d: 110. Distance PME-PME 160 µm, diameter PME 70 µm, distance PME- ALE ~30 µm; no trace of AME.
COLOR. Entire animal pale gray to whitish, legs slightly yellowish with brown patellae and tibia-metatarsus joints.
BODY. Habitus as in close congeners (cf.
Figs 122, 124
); ocular area barely elevated, each triad on very low hump; carapace without median furrow; clypeus unmodified; sternum as wide as long (0.40), unmodified.
CHELICERAE. As in
Figs 168–169
, apophyses near lamellae bipartite with small proximal part, with pair of barely visible lateral processes; without modified hairs; without stridulatory ridges.
PALPS. In general similar to
C. vermiformis
(cf. figs 139 and
140 in
Huber 2011
); trochanter apophysis as in
Fig. 167
; femur as in
Fig. 166
, distal process at 50% of femur length; tibia very slender (
Fig. 286
), length/diameter 0.44/0.15; procursus as in
Figs 164–165
, with simple ventro-distal flat sclerite; bulb length 0.30; embolus length 0.58; embolus tip with distinct fringes.
LEGS. Without spines and curved hairs; few vertical hairs; trichobothria on tibia 1 and tarsus 1 pseudosegments not seen.
Male
(variation). Tibia
1 in
2 other males: 5.5, 6.1. The male from
East Kalimantan
shares the distinctively slender palpal tibia but is significantly larger and therefore assigned tentatively (tibia 1 missing; palpal tibia length/ diameter 0.59/0.18; bulb length 0.38; embolus length 0.80).
Female
. In general similar to male, also eye triads at same distance. Tibia
1 in
3 females
: 5.5, 5.6, 5.8. Epigynum as in
Fig. 170
, weakly sclerotized triangular plate with small and slender anterior ‘knob’; internal genitalia as in
Fig. 171
, longer than wide, with oval pore plates close together.
Distribution
. Known from
Central Kalimantan
and (specimens assigned tentatively)
East Kalimantan
(
Fig. 283
).