Revision of fossil Bibionidae (Insecta: Diptera) from French Oligocene deposits Author Skartveit, John Author Nel, André text Zootaxa 2017 4225 1 1 83 journal article 37263 10.5281/zenodo.259087 05be601b-f9f1-4952-ac09-7012ef31aab0 1175-5326 259087 4676EE59-569B-4197-B108-9B32FD88E68E Penthetria lugens ( Oustalet, 1870 ) ( Figs. 14 – 15, 95 – 97) Protomyia lugens Oustalet, 1870 : 142 – 143. Plecia cf. lugens ( Oustalet, 1870 ) ; Théobald, 1937 : 227 . Penthetria lugens ( Oustalet, 1870 ) ; Handlirsch, 1908 : 959 . Material examined. Holotype (female) MNHN R 0 6666. Additional material: 2 females (MNHN B 24412, B 24518). 5 females (NHB F2208, F2230, F2323, F2410, F2413). 1 female (FSL 391923). Localities. Corent, Céreste, Kleinkembs The holotype specimen (fig. 12) is in very poor condition, wing venation and most details are impossible to see. Collomb et al . (2008) classified the type as Insecta, incerta sedis , though it is recognizable as a Plecia or Penthetria female. However, the diagnostic appendages on gonapophysis VIII are visible, confirming that the holotype is conspecific with Penthetria lugens sensu Théobald, 1937 . Diagnosis. A medium-sized, slender Penthetria . Body brownish, wings brown fumose. R2+3 slightly sinuous and oblique. On ventral side of abdomen gonapophysis VIII has a pair of digitiform appendages. Redescription. Male unknown Female (fig. 13) Total length 8.3 – 11.5 mm (N=4). Head (fig. 96): Length 0.93 – 1.11 mm (N=3), width 1.01 – 1.27 mm (N=3). Dark brown, probably black in life. Antennal flagellum probably 5- or 6-segmented. Thorax: Length 2.0 – 2.6 mm (N=4), width 2.0 mm (N=2). Dark brown, probably black in life. Haltere yellowish-brown. Wing (fig. 95): Length 8.6 – 9.4 mm (N=4), width 2.9 – 3.3 mm (N=4), length/width = 2.87 – 3.34 (N=4). Brown fumose, R-veins brown, posterior veins concolourous with membrane. Costa extends nearly to apex of M1. Humeral vein present, Subcosta fine and short, terminating well before crossvein R-M. Pterostigma indistinctive. R2+3 with a bend in basal third, otherwise nearly straight, oblique. R4+5 apically nearly straight. Posterior veins not possible to see in specimens at hand. Wing vein measurements in Table 1 . Legs: Brown, slender. Fore femur 2.0 mm (N=1), fore tibia 2.2 mm (N=1), hind femur 2.6 mm (N=1), hind tibia 2.7 mm (N=1). Abdomen: Length 6.1 – 7.6 mm (N=4), width 1.7 – 2.9 mm (N=3). Light brown, relatively long and narrow, spindle-shaped. Terminalia (fig. 97): Gonapophysis VIII with a digitiform appendage on each side. Cerci two-segmented, relatively long. FIGURES 95–97. Penthetria lugens (Oustalet, 1870) Fig. 95. Female, wing. NMB F2410. Fig. 96 . Female, head and thorax, dorsal. FSL 391923. Fig. 97 . Female, terminalia, ventral. NMB F2413.