Bryozoan faunas at the Tortonian-Messinian transition. A palaeoenvironmental case study from Crete Island, eastern Mediterranean Author Moissette, Pierre National & Kapodistrian University of Athens, Faculty of Geology and Geoenvironment, Department of Historical Geology and Paleontology 15784, Athens (Greece) and CR 2 P (CNRS, MNHN, UPMC, Sorbonne Université), Département Origines et Évolution, UMR 7207, Muséum national d’Histoire naturelle, case postale 38, 57 rue Cuvier, F- 75231 Paris cedex 05 (France) pmoissette@geol.uoa.gr Author Antonarakou, Assimina National & Kapodistrian University of Athens, Faculty of Geology and Geoenvironment Department of Historical Geology & Paleontology, 15784, Athens (Greece) aantonar@geol.uoa.gr Author Kontakiotis, George National & Kapodistrian University of Athens, Faculty of Geology and Geoenvironment Department of Historical Geology & Paleontology, 15784, Athens (Greece) gkontak@geol.uoa.gr Author Cornée, Jean-Jacques Géosciences Montpellier, Université de Montpellier Université des Antilles, CNRS, Pointe à Pitre, Guadeloupe, FWI (France) jean-jacques.cornee@gm.univ-montp2.fr Author Karakitsios, Vasileios National & Kapodistrian University of Athens, Faculty of Geology and Geoenvironment Department of Historical Geology & Paleontology, 15784, Athens (Greece) vkarak@geol.uoa.gr text Geodiversitas 2021 2021-12-16 43 26 1365 1400 journal article 20894 10.5252/geodiversitas2021v43a26 f8cedc0a-01dc-4bb0-855b-18064d237655 1638-9395 5796571 urn:lsid:zoobank.org:pub:14A6956D-54AD-48D2-9C5E-BA380EDACAA4 Cupuladria cf. canariensis ( Busk, 1859 ) ( Fig. 9A, B ) Cupularia canariensis Busk, 1859: 66 , pl. 23, figs 6-9. Cupuladria canariensis – Manzoni 1869 , p. 26, pl. 2, fig. 17; 1877: 72, pl. 17, figs 56a-c.— Cipolla 1921: 31 , pl. 2, figs 22-24.— Lagaaij 1952: 33 , pl. 2, figs 1a-b. — Buge 1957: 139 , pl. 9, fig. 5. — Annoscia 1963: 225 , pl. 9, fig. 1; pl. 10, fig. 1; pl. 11, figs 1a-b; pl. 12, figs 1a-b. — Cook 1965: 197 , text-figs 1a-f, pl. 1, fig. 1; pl. 3, fig. 4. — Prenant & Bobin 1966: 307 , figs 101-102. — Baluk & Radwansky 1984: 21, pl. 1, figs 1-4; pl. 8, figs 1-4. — Zabala & Maluquer 1988: 89 , fig. 112- 114. — Pouyet & Moissette 1992: 33 , pl. 3, figs 4-5. — Moissette et al. 1993: 92 , figs 4g-i. — Haddadi-Hamdane 1996: 65, pl. 5, figs 1, 4. — Marcopoulou-Diacantoni & Wuest 1999: 555, pl. 3, fig. 1. FIG. 9. — A , B , Cupuladria cf. canariensis ( Busk, 1859 ) ; A , frontal view of a whole colony, KER39: AMPG(IV) 3078a; B , dorsal view of a whole colony, KER39: AMPG(IV) 3078b; C , D , Discoporella reussiana ( Manzoni,1869 ) ; C , frontal view of a whole colony,KER30:AMPG(IV) 3102; D , dorsal view of a whole colony,KER30: AMPG(IV) 3101; E , Nellia tenella ( Lamarck, 1816 ) , detail of an internode showing two zooids in frontal view, KER17: AMPG(IV) 3150a; F -H , Canda rectangulata Udin, 1964 ; F , dorsal view of an internode fragment, FAN35: AMPG(IV) 3506a; G , frontal view of an internode fragment, FAN35: AMPG(IV) 3506b; H , Detail of the same fragment, FAN35: AMPG(IV) 3506b. Scale bars: A-D, 1 mm; E, H, 100 µm; F-G, 200 µm. OCCURRENCE. — Middle Miocene: France , Spain , Austria , Poland (Baluk & Radwansky 1984). Late Miocene: Germany , Italy , Crete ( Moissette et al. 1993 ). Pliocene: UK , Netherlands , Sicily ( Pouyet & Moissette 1992 ), Algeria (Haddadi-Hamdane 1996), Crete (Marcopoulou-Diacantoni & Wuest 1999). Pleistocene: Sicily ( Rosso 1987 ), Rhodes (Moissette 2012). Recent: eastern Pacific ( Ecuador to northern Mexico ), western and eastern Atlantic ( Brazil , Caribbean, Gabon , Azores , Madeira and Canary islands to southern Portugal ), southern Mediterranean ( Rosso & Di Martino 2016 ). This warm-water species lives on sandy, more or less muddy bottoms at depths between 50 and 300 m ( Prenant & Bobin 1966 ). But it has also been found in much shallower ( 5-50 m ) and much deeper waters (down to 860 m in the Sargasso Sea; Lagaaij 1963 ). Following Cadée (1979 , 1981 ), C. canariensis seems however mostly a western Atlantic species.Records with this name must consequently be carefully checked, eliminating possible misidentifications (A. Rosso, personal communication). REMARKS Due to possible confusion with two other species of the same genus ( C. biporosa (Canu & Bassler, 1923) and C. vindobonensis Baluk & Radwanski, 1984 ) a systematic revision is needed. Cadée (1979) also created a new subspecies, Cupuladria canariensis cavernosa , for Mio-Pliocene European specimens with intermediate characteristics between C. biporosa and C. canariensis ( Cadée 1979 , 1981 ).