Polyphyly of the traditional family Flabellinidae affects a major group of Nudibranchia: aeolidacean taxonomic reassessment with descriptions of several new families, genera, and species (Mollusca, Gastropoda) Author Korshunova, Tatiana Author Martynov, Alexander Author Bakken, Torkild Author Evertsen, Jussi Author Fletcher, Karin Author Mudianta, I Wayan Author Saito, Hiroshi Author Lundin, Kennet Author Michael Schroedl, Author Picton, Bernard text ZooKeys 2017 717 1 139 http://dx.doi.org/10.3897/zookeys.717.21885 journal article http://dx.doi.org/10.3897/zookeys.717.21885 1313-2970-717-1 C19B43B1B3214CB1B1B2A246CEAC56BC C19B43B1B3214CB1B1B2A246CEAC56BC Polaria gen. n. Figs 2, 7, 11 Type species. Coryphella polaris Volodchenko, 1946 Etymology. After the northern Polar region, the predominant area of distribution of this genus. Diagnosis. Body wide. Notal edge present, well-defined, continuous. Cerata not stalked, continuous. Rhinophores smooth to wrinkled, longer than oral tentacles. Anterior foot corners present. Anus pleuroproctic under the notal edge. Rachidian teeth with strong smooth cusp and distinct denticles. Lateral teeth strongly denticulated with considerably attenuated process basally. Single distal receptaculum seminis. Long vas deferens without separate granulated prostate. No penial collar. Penis elongated conical. Species included. Polaria polaris (Volodchenko, 1946), comb. n. (Figs 7, 11) (original description in Volodchenko 1946 ). Remarks. The type and single species of the genus Polaria , Coryphella polaris is the only available valid name ( Volodchenko 1946 ) for the species Goniaeolis typica M. Sars, 1861 as incorrectly identified by Bergh (1886) , Odhner (1907) , and Roginskaya (1987 , 1997 ). True Goniaeolis typica possesses a triserial radula but has no digestive gland branches or cnidosacs in the cerata and the shape of radular teeth is very different from any traditional flabellinid (M. Sars 1861 ; G.O. Sars 1872 ; Odhner 1922 ). The name Coryphella polaris Voldochenko, 1946 was therefore resurrected to avoid this misidentification ( Martynov 2006a ). The genus Polaria forms a separate clade within Paracoryphellidae according to the molecular phylogenetic analysis (Figs 1, 2). By combination of relatively long vas deferens without distinct prostate (Figs 7, 10H, I), retractable penis without external collar, and lateral teeth with strongly attenuated process (Fig. 10G), the genus Polaria morphologically differs from all other genera of the family Paracoryphellidae .