A new genus of air-breathing marine slugs from South-East Asia (Gastropoda, Pulmonata, Onchidiidae) Author Dayrat, Benoit Author Goulding, Tricia C. Author Khalil, Munawar Author Comendador, Joseph Author Xuan, Qu ảng Ngo Author Tan, Siong Kiat Author Tan, Shau Hwai text ZooKeys 2019 877 31 80 http://dx.doi.org/10.3897/zookeys.877.36698 journal article http://dx.doi.org/10.3897/zookeys.877.36698 1313-2970-877-31 ABBC811D2A6D46159ECF0F44933BFBE1 5BE094A8A1645289A9A3A154C4AF6293 Laspionchis bourkei lateriensis Dayrat & Goulding ssp. nov. Figs 17E , 18E, F, I , 19D , 21B , 22B , 23 A-D , 24E, G , 25 Holotype . INDONESIA • holotype, designated here, 17/16 mm [6064 H]; Ambon, Lateri; 03°38.261'S , 128°14.716'E ; 12 Feb. 2014; M Khalil and party leg.; st 128, mudflat next to small creek in the low intertidal of mangrove preserve; UMIZ 00115. Additional material examined. INDONESIA - Ambon • 5 specimens 12/7 mm [2743], 17/10 mm [2753], 15/12 mm [6061], 18/13 mm [6063], 15/12 mm [6065]; same collection data as for the holotype; UMIZ 00116. Distribution ( Fig. 7 ). Indonesia (Ambon). Habitat ( Fig. 17E ). Same as the entire species Laspionchis bourkei (see above). Etymology. The subspecies Laspionchis bourkei lateriensis is named after Lateri, in Ambon because the type locality is part of the preserved mangrove of Lateri. The name lateriensis is an adjective derived from Lateri and the suffix - ensis . Diagnosis (Table 3 ). Externally, the three subspecies of L. bourkei cannot be distinguished. Internally, L. bourkei lateriensis differs from L. bourkei bourkei but cannot be distinguished from L. bourkei matangensis . All three subspecies, however, are clearly delineated using molecular DNA sequences. Color and morphology of live animals ( Fig. 18E, F, I ). Identical to the species L. bourkei (see above). Digestive system ( Figs 19D , 25 ). Identical to the species L. bourkei (see above). Examples of radular formulae are presented in Table 4 . Radulae measure up to 1.7 mm in length. Figure 25. Radula, Laspionchis bourkei lateriensis , Indonesia, Ambon A-D holotype [6064 H] (UMIZ 00115) E, F [6063] (UMIZ 00116). A Rachidian and innermost lateral teeth B rachidian and innermost lateral teeth C lateral teeth D lateral teeth E lateral teeth (inferior view) F outermost lateral teeth. Scale bars: 10 μm ( A, F) , 20 μm ( B, C) , 30 μm ( D) , 8 μm ( E) . Abbreviations: bls basal lateral spine hlt hook of a lateral tooth. Reproductive system ( Fig. 21B ). Identical to the species L. bourkei (see above). Copulatory apparatus ( Figs 22B , 23 A-D , 24E, G ). Similar to the species L. bourkei (see above) acknowledging some minor variations: the length of the spine of the accessory penial gland ranges from 0.35 mm [6063] (UMIZ 00116) to 0.75 mm [6064 H] (UMIZ 00115), the retractor muscle is vestigial or absent, and penial hooks measure from 20 to 45 μm . Remarks. See above, the remarks on the species Laspionchis bourkei .