A redescription of TraCheloSaUrUS fiSCheri from the Buntsandstein (Middle Triassic) of Bernburg, Germany: the first European DinoCephaloSaUrUS-like marine reptile and its systematic implications for long-necked early
Author
Spiekman, Stephan N. F.
*
Author
Ezcurra, Martín D.
Author
Rytel, Adam
Author
Wang, Wei
Author
Mujal, Eudald
*
Author
Buchwitz, Michael
Author
Schoch, Rainer R.
*
text
Swiss Journal of Palaeontology
2024
10
2024-03-15
143
1
1
33
http://dx.doi.org/10.1186/s13358-024-00309-6
journal article
10.1186/s13358-024-00309-6
1664-2384
12004587
Trachelosauridae
Abel, 1919
1
(= Dinocephalosauridae
Spiekman, Fraser and Scheyer, 2021
)
Phylocode registration number.
Trachelosauridae
is identified in the international clade names repository as registration number 1028.
Phylogenetic definition.
The definition of
Trachelosauridae
is modified from the definition for Dinocephalosauridae (
Spiekman et al., 2021b
) as follows: the most inclusive clade containing
Trachelosaurus fischeri
Broili, 1918
and
Dinocephalosaurus orientalis
Li, 2003
but not
Tanystropheus longobardicus
(
Bassani, 1886
)
,
Macrocnemus bassanii
Nopcsa, 1931
,
Protorosaurus speneri
von
Meyer, 1832
, or
Prolacerta broomi
Parrington, 1935
. This is a maximum clade definition.
Reference phylogeny.
Phylogenetic hypotheses recovered in this paper.
Composition.
The composition is based on our reference phylogenies.
Trachelosauridae
includes the following nominal species:
Dinocephalosaurus orientalis
,
Trachelosaurus fischeri
,
Austronaga
minuta
,
Pectodens zhenyuensis
, and ambiguously
Gracilicollum latens
, and
Fuyuansaurus acutirostris
.
Diagnosis.
Trachelosauridae
is a clade that is defined among other archosauromorphs by possessing the following unique combination of character states: external naris positioned far from the anterior end of the premaxilla, resulting in an anteroposteriorly deep base of the prenarial process; jugal without posterior process; cervical column composed of ten or more vertebrae (also present in some deeply nested tanystropheids); axis with postzygapophysis confluent with and not protruding posteriorly from the base of the neural spine in lateral view (also present in
Tanystropheus
spp.
and
Amotosaurus rotfeldensis
); mid-dorsal vertebrae with very wide and ‘wing-like’ transverse processes; presence of holocephalous anterior dorsal ribs (also present in some tanystropheids); metatarsal V without a hook-shaped proximal end.