Lance lacewings of the world (Neuroptera: Archeosmylidae, Osmylidae, Saucrosmylidae): review of living and fossil genera
Author
Winterton, Shaun L.
Author
Martins, Caleb Califre
Author
Makarkin, Vladimir
Author
Ardila-Camacho, Adrian
Author
Wang, Yongjie
text
Zootaxa
2019
2019-04-09
4581
1
1
99
journal article
27348
10.11646/zootaxa.4581.1.1
5b259853-01f6-4307-8bef-58c98ec73bd3
1175-5326
2633842
20A9776D-AE5F-41BC-A35B-0C5E42EDFE48
Lithosmylus
Carpenter, 1943
Type
species.
Osmylus columbianus
Cockerell, 1908
: 758
(by original designation).
Diagnosis.
Wings oval; FW costal area broad basally, subcostal veinlets closely spaced and mostly shallowly forked in both wings, veinlets simple at wing base and distally; first branch of RP originating near origin of RP on RA; FW RP with 16 branches, HW with 17 branches; both wings with outer gradate series well developed, inner gradate series absent, radial crossveins numerous in basal half of wing, slightly more irregularly arranged in HW; M forked slightly distal to origin of RP1; MA and MP with 2–4 branches distally in both wings; CuA and CuP strongly pectinately branched in both wings with shallow end-twigging very close to wing margin, FW with six branches in CuA and nine branches in CuP, HW with seven branches in CuA and 12 branches in CuP; FW A1 with seven pectinate branches, HW A1 with four pectinate branches; FW A2 with four pectinate branches, two in HW; A3 simple in both wings.
Comments.
The venation of this fossil genus is relatively typical for
Osmylinae
and we agree with its placement in this subfamily by
Lambkin (1987)
and
Wang
et al.
(2011a)
, although considering the wing venation similarity,
Carpenter (1943)
unsurprisingly regarded it as belonging to
Kempyninae
. Only the wings are known.