The genus Argopistes Motschulsky from Japan and Taiwan, with descriptions of three new species from Taiwan (Coleoptera, Chrysomelidae, Galerucinae, Alticini) Author Lee, Chi-Feng 0000-0003-1996-0557 Applied Zoology Division, Taiwan Agricultural Research Institute, Taichung 413, Taiwan Author Chiang, Ming-Yao 0000-0002-1514-7086 Applied Zoology Division, Taiwan Agricultural Research Institute, Taichung 413, Taiwan Author Suenaga, Haruki Nakashima, 108 - 11, Kurashiki-shi, Okayama, 710 - 0803 Japan text ZooKeys 2024 2024-10-15 1215 151 183 journal article 10.3897/zookeys.1215.134871 C57CB315-F15F-4D98-868A-EFEA22BC64A3 Argopistes tsekooni Chen, 1934 Figs 8 A – C , 9 Argopistes tsekooni Chen, 1934 b: 316 ( China : Shanghai , Hangchow); Csiki 1940: 525 (catalogue); Chûjô and Kimoto 1961: 174 ( China , Japan ); Gressitt and Kimoto 1963: 813 ( China : Jiangsu ); Kimoto 1965: 437 (redescription); Lee and An 2001: 183 ( South Korea ); Lee and Cho 2006: 91 (host plant); Takizwa, 2012: 38 (faunistics); Cho and An 2020: 15 ( North Korea ); Won et al. 2023: 9 ( South Korea : Ulleungdo). Argopistes biplagiatus : Baly 1874: 202 (misidentification). Type material examined. One syntype • (sex undetermined, IZAS ): “ 浙江 (= Zhejiang ): 杭州 (= Hangchow) / 1934. [h] / 中国科學院 (= Chinese Academy of Sciences) [h, p] // 害水蜡樹 (attacking Ligustrum obtusifolium ) [h, w] // Argopistes / tsekooni / Chen [h, w] ”. Although this specimen does not bear any type label, it should be regarded as type specimen since it fit the original description. Additional material examined. Japan . 1 ♀ ( NHMUK ): “ Argopistes / biplagiatus / Motsch / Japan [h, w] // Baly Coll. [p, w] ”; Honshu. Shizuoka : • 1 ♂ ( SEHU ), Tagata-gun , Tohi , 4. V. 1985 , leg. Y. Komiya ; Tokyo : • 1 ♂ ( HAPC ), Komae-shi , Komai-machi , 10. VI. 2021 , leg. R. Seki ; Yamaguchi : • 1 ♂ ( NHMUK ); Kyushu. Fukuoka : • 1 ♀ ( HAPC ), Fukuoka-shi , Higashi-ku , Shimobaru (alt. 100–360 m ), 27. V. 2009 , leg. S. Sejima ; • 1 ♀ ( NHMUK ), Mt. Mikazuki , 2. V. 1954 , leg. K. Morimoto ; Nagasaki : • 1 ♂ , 2 ♀ ( SEHU ), Sasebo-shi , Mt. Yahirodake , 14. IV. 1981 , leg. J. Okuma ; • 1 ♂ ( SEHU ), same locality but with “ 21. IV. 1981 ”; Oita : • 2 ♂ , 3 ♀ ( HAPC ), Hita-shi , Miwa , Chikura , 11. IV. 2016 , leg. S. Sasaki. Diagnosis. Adults of A. tsekooni are recognized easily by their small body sizes (< 3.5 mm ;> 3.5 mm in others except A. unicolor ), elongate ovate body shapes (elytra 1.2 × longer than wide; but 1.1 × longer than wide in others), and the combined red spots on elytra (usually separate red spots on the elytra in others); additionally, most genitalic characters are unique, such as the tube-like apex of the aedeagus (Fig. 9 C ); few setae on apical margin of abdominal ventrite VIII in females (Fig. 9 E ); and transverse gonocoxae with dense, long setae on the widely rounded apical margin (Fig. 9 G ). Redescription. Length 2.8–3.2 mm , width 2.1–2.4 mm . Color (Fig. 8 A – C ) blackish brown, elytron with one transverse orange area at basal 1 / 3, and narrowed towards suture; tarsi and front tibiae yellow; antennae dark brown but seven basal antennomeres yellow. Pronotum broad, convex, lateral margin narrowly explanate; 2.0–2.1 × wider than long, disc with dense coarse punctures; lateral margin rounded, anterior margin strongly concave, posterior margin moderately convex. Elytra elongate oval, 1.2 × longer than wide, disc with confused, dense, coarse punctures. Habitus of Argopistes species A A. tsekooni Chen , male, dorsal view B ditto, ventral view C ditto, lateral view D A. unicolor Jacoby , female, dorsal view E ditto, ventral view F ditto, lateral view. Argopistes tsekooni Chen A antenna, male B antenna, female C aedeagus, dorsal view D aedeagus, lateral view E abdominal ventrite VIII, female F spermatheca G gonocoxae. Male. Antenna filiform (Fig. 9 A ), antennomere I much longer than others, approximate ratios of length of antennomeres I – XI 1.0: 0.4: 0.3: 0.5: 0.5: 0.5: 0.5: 0.5: 0.5: 0.5: 0.7; approximate ratios of length to width of antennomeres I – XI 3.1: 1.8: 2.0: 2.5: 2.3: 1.9: 1.8: 2.1: 2.1: 2.2: 2.9. Aedeagus (Fig. 9 C, D ) gradually widened from basal 1 / 9–1 / 2, then gradually narrowed to basal 1 / 2, strongly widened posterior – basal 1 / 2; anterior opening large, ~ 0.39 as long as aedeagus, from apex to apical 2 / 5; tectum composed of one pair of sclerotized processes, long, ~ 0.85 as long as anterior opening, wide and slightly curved from basal 2 / 4 to apex in lateral view, recurved near apex; endophallic sclerite laterally flattened, with base twisted. Female. Antenna (Fig. 9 B ) similar to males, but antennomeres VII – X wider, ratios of length of antennomeres I – XI 1.0: 0.4: 0.3: 0.4: 0.4: 0.4: 0.4: 0.4: 0.4: 0.4: 0.6; ratios of length to width of antennomeres I – XI 3.8: 1.8: 2.0: 1.8: 1.9: 1.9: 1.6: 1.5: 1.5: 1.5: 2.4. Ventrite VIII (Fig. 9 E ) weakly sclerotized, only part of apical margin well sclerotized, with several setae along apical margin, spiculum long and base wider. Spermathecal receptaculum (Fig. 9 F ) longer than pump, moderately swollen, curved in lateral view; pump emarginated at inner side of base; spermathecal duct with long basal part, ramus truncate. Gonocoxae (Fig. 9 G ) wide and separated, base membranous, each gonocoxa asymmetric, apically narrowed from near base, with sparse setae along apical areas, setae longer at apical 1 / 2. Color variation. One male has a black body and lacks transparent spots on elytra. Another male has an entire yellowish-brown body. Host plants. Oleaceae : Ligustrum obtusifolium ( Chûjô & Kimoto, 1961 ) ; Syringa oblata Lindl. , L. japonicum , L. licidum , and L. sinense ( Zhang et al. 2008 b ) . Biology. The biology and life history of A. tsekooni were studied under laboratory and outdoor conditions in Huangshan City of Anhui Province , China ( Zhang et al. 2009 ). Argopistes tsekooni overwintered as adults and had three overlapping generations in Anhui Province . Distribution. China , Japan (Honshu, Kyushu, the Goto Isls., Hirado-jima Is. Tsushima Is.), North Korea , South Korea .