Review of the North American species of Marimatha Walker with descriptions of three new species (Lepidoptera, Noctuidae, Eustrotiinae) and the description of Pseudomarimatha flava (Noctuinae, Elaphriini), a new genus and species confused with Marimatha Author Ferris, Clifford , Laramie, United States of America Author Lafontaine, Donald Agriculture and Agri-Food Canada, Ottawa, Canada text ZooKeys 2010 2010-03-18 39 39 117 135 journal article 10.3897/zookeys.39.424 4e1adc72-60c9-4846-af21-ed080ed81401 1313–2970 576637 66F92D10-9BCF-4B7A-AAD9-AF6042E8CEA0 Marimatha piscimala Ferris & Lafontaine , sp. n. urn:lsid:zoobank.org:act: F610BE0F-C8CF-4BA7-9C3D-BB88F91EB1DF Figs 9, 10, 15, 22 Type material. Holotype ♁. Texas . , Kimble Co. , Fred Coleman Ranch , 12 mi S Junction , 16 June 1972 , B. Wright ; Genitalia CNC slide # 15033 m . CNC . Paratypes : 22 ♁, 17 ♀ . USA . Arizona . Cochise Co. , Ash Canyon , Huachuca Mts. , 5170’, 20 July 2006 ( 1 ♀ ), 4 Aug. 2009 ( 1 ♀ ), C. D. Ferris ; Pima Co. , Baboquivari Mts. , Brown Canyon , 3880’, 18–19 Aug. 2006 , C. D. Ferris (6 ♁, 3 ♀ ) ; Pima Co. , Baboquivari Mts. , Brown ( Bear ) Canyon , 5300–5900’, 2 Aug. 2008 , C. Schmidt & L. Crabo , mesquite/riparian (1 ♁) ; Santa Cruz Co. , Madera Canyon 4880’, 13 July – 28 Aug. 1959 , J. G. Franclemont (3 ♁, 3 ♀ ) ; Santa Cruz Co. , Peña Blanca 3950’, 7 Aug. 1959 (1 ♁), 23 Aug. 1960 (1 ♁), J. G. Franclemont ; Santa Cruz Co. , Patagonia , 4050’, 8 Aug. 2009 , C. D. Ferris (1 ♁) . Texas . Cameron Co. , Audubon Sabal Palm Grove , 11 Oct. 1996 , C. Bordelon ( 1 ♀ ), 20 April 1997 E. C. Knudson ( 1 ♀ ) ; Comal-Hayes Counties , El Rancho , 29 Aug. 1975 , A . & M. E. Blanchard (1 ♁) ; Hidalgo Co. , Bentsen State Park , 18–19 April 1997 , E. C. Knudson ( 1 ♀ ) ; Hidalgo Co. , Mission , 12 Oct. 2008 , C. W. Bordelon (1 ♁) ; Jeff Davis Co. , Ft Davis , Jeff Mts. State Park , 11–13 Sept. 2008 , Knudson / Bordelon (1 ♁, 1 ♀ ) ; Kerr Co. , Kerrville State Park , 19 Aug . Figures Į8 – 22. Marimatha female genitalia. Į 8 M . nigrofimbria Į 9 M . squala 20 M . quadrata 2Į M . tripuncta 22 M . piscimala . 1980, E. C. Knudson ( 1 ♀ ); Starr Co., Roma, 6,7 Oct. 1996 , E. C. Knudson ( 1 ♀ ). Mexico . Chiapas : 5 mi NW Pijijiapan , Hwy. 200, 31 May 1969 , A . Mutuura (5 ♁, 1 ♀ ) ; Chiapas , 15 May, R . Muller ( 1 ♀ ) ; San Luis Potosi , El Naranjo , El Salto , 29 June 1965 , P. Spangler ( 1 ♀ ) . Panama . La Chorrera , 12 May , August Busck (1 ♁). Paratypes deposited in CDF , CNC , CUIC , JBW , TLSC , and USNM . Other material examined. USA . Arizona . Cochise Co., Ash Canyon, Huachuca Mts, 31°23.17'N 110°14.281'W , 5170’, 4 August 2009 , C. D. Ferris. Santa Cruz Co., Patagonia (in town), 31°32.62'N 110°45.31'W , 4050’, 8 August 2009 , C. D. Ferris. Costa Rica . Guanacaste , Area de Conservacion Guanacaste , Sector Santa Elena, La Angostura, 26 May 2006 ( 1 ♀ ), 24 July 2006 (1 ♁), F. Quesada & R. Franco. Etymology . The species name is derived from the Latin for fish ( Piscis ) and jaw ( mala ) and refers to the fish-jaw like appearance of the male valva. Diagnosis . Head with frontal protuberance. Male genitalia: valva broad, ventral margin even; dorsal margin slightly concave to middle, then tapering to apex with multiple narrow dentate projections from dorsal margin; vesica with a long apical cornutus and postmedial cluster of cornuti with mixture of two large and many small cornuti. Female genitalia: ostial plate heart shaped with cleft in middle of posterior margin extending almost to base of plate. Description . Adult: Head – Frontal protuberance shallow crater-like prolate ellipsoidal. Thorax – Yellow with brown flecks. Legs : yellow, flecked with brown scales, more heavily on forelegs and middle legs, and in females. Wings (both sexes): DFW golden yellow with dark discal spot and additional dark spot in fold at ¼ from wing base; often with a smaller third spot below costa at 1/4 from base; a thin orange-brown postmedial line deeply curving around discal spot in some males and most females; specimens from southeastern Texas , and less commonly those from elsewhere, may have increased dark shading on the postmedial line and sometimes a longitudinal streak along the middle of the wing like that often present in M . tripuncta . DHW pale fuscous. VFW yellow, heavily infused with brown scales with well-defined pale band along inner margin. VHW yellow, moderately flecked with brown scales, indistinct brown marginal line; in some specimens a suggestion of a brown discal spot and postmedial line. Male genitalia – Uncus : decurved, slender, slightly spatulate tip. Valvae : symmetrical, broad basally, ventral margin even; dorsal margin slightly concave to middle, then tapering sharply to apex with multiple (7 typically) narrow dentate projections from dorsal edge; apex pointed, corona absent. Aedeagus : smoothly sclerotized; vesica with one large apical cornutus and postmedial cluster of smaller cornuti. Female genitalia – Corpus bursae elongate without surface features; ductus seminalis on right side about ⅓ from anterior end. Anterior part of ductus bursae broad and lightly sclerotized, ductus constricted and elbowed at 1/3 from anterior end with posterior 2/3 narrower and increasingly heavily sclerotized. Ostial plate heart shaped with cleft in middle of posterior margin extending almost to base of plate; when apex of plate brushed in undissected specimens protruding sides of plate project posterolaterally. Distribution and biology. Southeastern Texas to Arizona southward to Panama . Flight season mid-April to mid-October, probably several generations.