Twenty-one new species of Omaliini from the Papuan Region (Coleoptera: Staphylinidae: Omaliinae), with diagnostic and faunistic notes on some species of the genus Paraphloeostiba Steel, 1960 Author Shavrin, Alexey V. text Zootaxa 2024 2024-03-14 5424 3 251 307 http://dx.doi.org/10.11646/zootaxa.5424.3.1 journal article 10.11646/zootaxa.5424.3.1 1175-5326 10821214 DE26201A-252D-40D3-A5CF-FCCC6B3C2D01 Paraphloeostiba iriana sp. n. ( Figs 40 , 64–67 ) Type material. Holotype ♂ (dissected): ‘ IRIAN JAYA : Paniai Prov. | Bilogay, 2100-2200m | 29.12.1995 | leg. A. Riedel’ <printed>, ‘ HOLOTYPE | Paraphloeostiba | iriana sp. n. | Shavrin A. V . des. 2024’ <red, printed> ( NMW ). Paratypes : 2 ♂♂ : same data as the holotype, with additional red printed label: ‘ PARATYPE | Paraphloeostiba | iriana sp. n. | Shavrin A. V . des. 2024’ <red, printed> ( NMW ). Description. Measurements (n=3): HW: 0.35–0.37; HL: 0.20–0.21; OL: 0.12; TL: 0.02; AL ( holotype ): 0.57; PL: 0.24–0.27; PWmax: 0.41–0.47; PWmin: 0.38–0.43; ESL: 0.48–0.50; EW: 0.56–0.57; MTbL ( holotype ): 0.27; MTrL ( holotype ): 0.17 (MTrL 1–4: 0.07; MTrL 5: 0.10); AW: 0.55–0.57; AedL: 0.37; BL: 1.72–1.96 ( holotype : 1.92). Habitus as in Fig. 40 . Body and antennomeres 6–11 brown, with slightly paler laterobasal and basal margins of pronotum and middle portions of each elytron; mouthparts, antennomeres 1–5 and legs yellow. Head, neck, pronotum and scutellum without visible punctation; elytra with very fine and sparse indistinct punctation, slightly denser in basal portion and invisible along suture; abdominal tergites with very fine and sparse punctation, indistinct in middle portions of abdominal tergites IV–VI. Head with dense and coarse microsculpture, transverse on vertex and isodiametric in middle; pronotum with very dense isodiametric microsculpture; microsculpture similar to that in pronotum, but slightly finer and longer; abdominal tergites with very dense, moderately coarse and isodiametric microreticulation. Head with distinct moderately deep, wide and oval anteriomedian depressions, with short anteocellar foveae, about twice as long as diameter of each ocellus; poctocular carina very indistinct. Ocelli moderately large, located slightly below level of posterior margins of eyes; distance between ocelli about 1.4 times as long as distance between ocellus and posterior margin of eyes. Antennomere 4 about as long as wide, disitnctly shorter than 3, 5 slightly shorter and indistinctly broader than 4, 6 significantly broader than 5, 7–9 indistinctly longer than 6, 10 slightly longer and indistinctly broader than 9, apical antennomere about 1.6 times as long preapical segment, from middle gradually narrowed apicad toward rounded apex. FIGURES 64–67. Paraphloeostiba iriana (holotype): 64–aedeagus, parameral view, 65–aedeagus, lateral view, 66–male abdominal tergite VIII, 67–male abdominal sternite VIII. Scale bar: 0.1 mm. Pronotum 1.7 times as broad as long, 1.1–1.2 times as broad as head, widest in about middle, distinctly more narrowed posteriad than anteriad; middle surface with moderately wide shallow impressions, broadened toward mediobasal margin; laterobasal impressions wide and shallow. Elytra about twice as long as pronotum. Metatarsi 1.5 times as long as metatibia. Abdomen about as broad as elytra. Male. Posterior margin of abdominal tergite VIII indistinctly sinuate ( Fig. 66 ). Posterior margin of abdominal sternite VIII widely and deeply concave ( Fig. 67 ). Aedeagus with wide basal portion, gradually narrowed toward rounded apex; parameres slightly exceeding apex of median lobe, moderately narrow, with two long apical and numerous long setae along inner margin of each paramere; internal sac wide and moderately long, with several short and elongate sclerotized structures in basal portion ( Fig. 64 ). Lateral aspect of the aedeagus as in Fig. 65 . Female unknown. Comparative notes. Based on the general structure of the aedeagus, P . iriana sp. n. is similar to P . guadalcanalensis sp. n. , P. riedeli sp. n. and P. conjuncta . Based on the presence of two distinct longitudinal impressions on the pronotum, it is more similar to P . guadalcanalensis sp. n. , from which it can be distinguished by the smaller body, slightly longer parameres with distinctly narrower apical portions. It can be distinguished from P. conjuncta and P. riedeli sp. n. by the broader apical portion of the median lobe. From both these species it can be distinguished by the slightly longer parameres. Additionally, it can be distinguished from P. conjuncta and P. riedeli sp. n. by the finer punctation of the head, the presence of longitudinal impressions in the middle of the pronotum, and details of the morphology of the aedeagus. From all these species it differs by the details of the external and internal morphology of the aedeagus, and the different shape of the female accesssory sclerite. Distribution. Paraphloeostiba iriana sp. n. is known only from the type locality in Paniai Province of Western New Guinea , Indonesia . Etymology. The specific epithet is the Latinized adjective derived from the name of the province of Irian Barat before being renamed Irian Jaya (literally “Glorious Irian”). Bionomics. Specimens were collected at elevations from 2100 to 2200 m a.s.l. The detailed bionomical data are unknown.