Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae) Author Bouzan, Rodrigo Salvador Pós-graduação em Zoologia, Instituto de Biociências, Universidade de São Paulo, São Paulo, Brasil. Author Iniesta, Luiz Felipe M. 0000-0002-0529-4162 Pós-graduação em Zoologia, Instituto de Biociências, Universidade de São Paulo, São Paulo, Brasil. & luiz-moretti @ hotmail. com; https: // orcid. org / 0000 - 0002 - 0529 - 4162 luiz-moretti@hotmail.com Author Brescovit, Antonio Domingos 0000-0002-1511-5324 antonio. brescovit @ butantan. gov. br; https: // orcid. org / 0000 - 0002 - 1511 - 5324 antonio.brescovit@butantan.gov.br text Zootaxa 2021 2021-05-14 4970 2 201 256 journal article 6318 10.11646/zootaxa.4970.2.1 89d3c795-198b-4162-8903-9c6cf3f1e665 1175-5326 4761544 6DBA0C02-9E52-4FD0-8B48-8C7B405713A8 Angelodesmus Schubart, 1962 Angelodesmus Schubart, 1962: 258 . Type species: A. costalimai Schubart, 1962 , by original designation; Hoffman, 1976: 173 ; Hoffman, 1980: 151 ; Shelley et al. 2000: 84 Diagnosis . Males of Angelodesmus differ from those of other Arthrosolaenomeridini genera by the combination of the following characters: stipes of the gnathochilarium with lateral folds; gonocoxae acuminate in ectal view ( Fig. 19F ); dorso-basal lobe in the prefemoral process absent ( Fig. 19D ); apex of the solenomere with the terminal zone marked by a stretch mark ( Figs 19D, F , arrows). Description. General characters : Coloration (specimen long preserved in 70% ethanol) whitish yellow, paranota tip and posterior border of the metaterga slightly whitish. Gnathochilarium: lingual plate covered completely by sparsely setae; mentum only centrally covered by setae, with smooth edges and stipes completely covered by sparsely setae and positioned mainly in the median region, presenting lateral folds. Body rings: cuticle slightly rough; alignment of paranota in posterior view straight; sternite of the fourth segment with a pair of projections, sternites of the fifth and sixth segments with two pairs of intumescences also covered by setae, the posterior pair less conspicuous; leg pair of the third body ring missing. Sternite of sixth body ring with two pairs of ventral projections; seventh body ring without projections and post-gonopodal sternites with two pairs of rounded ventral projections ( Fig. 30D ). Paranota form: round ( Fig. 19B ). Legs: without modifications only a slightly dorsal lobe on prefemur ( Fig. 31D ). Male characters : Leg pair of the third body ring missing, as well as the genital papilla. Posterior edge of the gonopod aperture without projections, excavated. Gonopods: gonocoxae equivalent to about half the length of the telopodite, cylindrical and acuminated in ectal view, without a spiniform process ( Fig. 19F ). Prefemoral process: elongate and thin, without a dorso-basal process ( Fig. 19D ). Cingulum in a middle position ( Fig. 19F ). Solenomere also long and thin with apex ascending ( Fig. 19D ). Female characters : Unknown. Distribution. State of Goiás , Brazil ( Fig. 28 ). Composition. Now considered as monotypic: Angelodesmus costalimai Schubart, 1962 .