Taxonomy of the leaf beetle genus Exomis Weise (Coleoptera: Chrysomelidae: Clytrini) with description of six new species from China
Author
Wang, Feng-Yan
Author
Zhou, Hong-Zhang
text
Zootaxa
2020
2020-03-06
4748
2
351
364
journal article
10.11646/zootaxa.4748.2.7
ae7f2b81-fd03-4ded-b33d-9f086dd3a0c8
1175-5326
3698881
1110012B-A68A-449D-BA03-AEB0DE911BFB
Genus
Exomis
Weise, 1889
Weise, 1889: 563
, 577;
Jacoby & Clavareau, 1906: 48
(subg. of
Gynandrophthalma
);
Clavareau, 1913: 67
(subg. of
Cyaniris
);
Gressitt, 1942: 328
;
Gressitt & Kimoto, 1961: 93
;
Seeno & Wilcox 1982: 33
(synonym of
Smaragdina
);
Regalin & Medvedev 2010a: 78
(resurrected from synonymy with
Smaragdina
),
Regalin & Medvedev, 2010b: 569
(catalogue).
Type
species:
Exomis peplopteroides
Weise, 1889
, fixed by monotypy.
Diagnosis:
The genus
Exomis
Weise, 1889
, can be recognized amongst other
Clytrini
by the following characteristics: body cylindrical, elongate; antennae short, 1
st
antennomere strongly swollen, 2
nd
and 3
rd
antennomeres small; Pronotum convex, as wide as or wider than elytra in males. Scutellum elevated toward posterior angle and above elytral plane. Epipleura pubescent. Legs robust, tarsi short, first tarsomere longer than second or third. Pygidium not exposed in dorsal view.
The aedeagi in this genus are different from other genera: elongate, tegmen situated at apical part of the median lobe, basal half of median lobe behind tegmen feebly sclerotized or membranous. The characteristics of the aedeagi are not as reliable as in other genera of the tribe
Clytrini
for species recognition. The female spermatheca and kotpresse sclerites are useful species diagnosing structures. The following determination key to species is a balanced one, and easy to use. For exact species recognition it is helpful to consider both genital characters of both sexes and external morphological structures.
Key to Chinese species of the genus
Exomis
Weise, 1889
1. Elytra pubescent...................................................................................... 2
- Elytra glabrous, at most lateral sides with a short row of setae.................................................. 3
2. Pronotum pubescent (male body length
5.1 mm
; female unknown)..............................
E. pubescens
sp. nov.
- Pronotum glabrous (male body length
6.8–6.9 mm
; female unknown)...........................
E. pubipennis
sp. nov.
3. Elytra almost entirely metallic green or blue, with a fulvous apical spot (5.4–7.0 mm).................
E. viridis
sp. nov.
- Elytra without metallic tinge............................................................................ 4
4. Elytra completely fulvous, without dark markings, at most an indistinct spot at humerus............................. 5
- Elytra black with fulvous markings, or fulvous with black markings............................................. 7
5. Abdomen and thorax entirely dark........................................................................ 6
- Venter with episternum and epimeron fulvous, body length
3.7–5.3 mm
.........................
E. deqinensis
sp. nov.
6. Legs entirely black........................................................................
E. degenevei
Pic
- Femora except for black apex orange, body length
4.6–5.4 mm
..........
E. oblongum
(Lopatin & Konstantinov)
com. nov
.
7. Elytra black, each elytron with a round fulvous spot at apex, pronotum fulvous with a central black spot extending both to anterior and posterior margins, body length
4.6–5.7 mm
).......................................
E. huangi
sp. nov.
- Elytra fulvous with black humeral spot or several black markings............................................... 8
8 Elytra fulvous with several black markings (
5.3–6.7 mm
)...................................
E. peplopteroides
Weise
- Each elytron with a black humeral spot, legs mostly fulvous (4.6–6.0 mm)............................
E. tanae
sp. nov.